John R. “Jack” Horner - All Publications - 1978 to 2016
Peer Reviewed Articles (115): (15 in Science & Nature)
Schram, F. R. and Horner, J. R. 1978. Crustacea of the Mississippian Bear Gulch Limestone of central Montana. Journal of Paleontology 52(2):394–406.
Baird, D. and Horner, J. R. 1979. Cretaceous dinosaurs of North Carolina. Brimleyana 2:1–28.
Horner, J. R. 1979a. Upper Cretaceous dinosaurs from the Bearpaw Shale (marine) of south-central Montana with a checklist of Upper Cretaceous dinosaur remains from marine sediments in North America. Journal of Paleontology 53(3):566–577.
Horner, J. R. and Makela, R. 1979. Nest of juveniles provides evidence of family structure among dinosaurs. Nature 282:296–298.
Horner, J. R. 1982. Evidence of colonial nesting and 'site fidelity' among ornithischian dinosaurs. Nature 297:675–676.
Horner, J. R. 1983. Cranial osteology and morphology of the type specimen of Maiasaura peeblesorum (Ornithischia: Hadrosauridae), with discussion of its phylogenetic position. Journal of Vertebrate Paleontology 3(1):29–38.
Horner, J. R. 1984a. A "segmented" epidermal tail frill in a species of hadrosaurian dinosaur. Journal of Paleontology 58(1):270–271.
Horner, J. R. 1984c. Three ecologically distinct vertebrate faunal communities from the Late Cretaceous Two Medicine Formation of Montana, with discussion of evolutionary pressures induced by seaway fluctuations. Montana Geological Society Guidebook 1984:299–303.
Horner, J. R. 1985. Stratigraphic position of the Bear Gulch Limestone (Lower Carboniferous) of central Montana. C.r. IX Congr. Int. Strat. Geol. Carbonifére 5:427–436.
Horner, J. R. and Lund, R. 1985. Biotic distribution and diversity in the Bear Gulch Limestone of central Montana. C.r. IX Congr. Int. Strat. Geol. Carbonifére 5:437–442.
Weishampel, D. B. and Horner, J. R. 1986. The hadrosaurid dinosaurs from the Iren Dabasu fauna (Peoples Republic of China, Late Cretaceous). Journal of Vertebrate Paleontology 6(1):38–45.
Horner, J. R. 1987. "Ecological and behavioral implications derived from a dinosaur nesting site"; pp. 51–63 in S. J. Czerkas and E. C. Olson (eds.), Dinosaurs Past and Present, Vol. II. University of Washington Press, Seattle, Washington.
Weishampel, D. B. and Horner, J. R. 1987. "Dinosaurs, habitat bottlenecks, and the St. Mary River Formation"; pp. 224–229 in P. Currie and E. H. Koster (eds.), Fourth Symposium on Mesozoic Terrestrial Ecosystems. Occasional Paper of the Tyrrell Museum of Palaeontology, Drumheller, Alberta, Canada.
Horner, J. R. 1988b. A new hadrosaur (Reptilia: Ornithischia) from the Upper Cretaceous Judith River Formation of Montana. Journal of Vertebrate Paleontology 8(3):314–321.
Horner, J. R. and Weishampel, D. B. 1988. A comparative embryological study of two ornithischian dinosaurs. Nature 332:256–257.
Horner, J. R. 1989a. The Mesozoic terrestrial ecosystems of Montana. Montana Geological Society Guidebook 1989:153–162.
Horner, J. R. 1990. "Evidence of diphyletic origination of the hadrosaurian (Reptilia: Ornithischia) Dinosaurs" - Chap. 13; pp. 179–187 in K. Carpenter and P. Currie (eds.), Dinosaur Systematics: Perspectives and Approaches. Cambridge University Press, New York, New York.
Weishampel, D. B. and Horner, J. R. 1990. "Hadrosauridae" - Chap. 26; pp. 534–561 in D. B. Weishampel, P. Dodson and H. Osmolska (eds.), The Dinosauria. University of California Press, Berkeley, California.
Horner, J. R. 1992a. Cranial morphology of Prosaurolophus (Ornithischia: Hadrosauridae) with descriptions of two new hadrosaurid species and an evaluation of hadrosaurid phylogenetic relationships. Museum of the Rockies Occasional Paper #2:1–119, Bozeman, Montana.
Horner, J. R. Varricchio, D. J. and Goodwin, M. B. 1992. Marine transgressions and the evolution of Cretaceous dinosaurs. Nature 358:59–61.
Barreto, C., Albrecht, R. M., Bjorling, D. E., Horner, J. R. and Wilsman, N. J. 1993. Evidence of the growth plate and the growth of long bones in juvenile dinosaurs. Science 262:2020–2023.
Rogers, R. R., Swisher, C. C., Horner, J. R. 1993. Ar40/Ar39 age and correlation of the nonmarine Two Medicine Formation (Upper Cretaceous), northwestern Montana, USA. Canadian Journal of Earth Sciences 30:1066–1075.
Varricchio, D. J. and Horner, J. R. 1993. Hadrosaurid and Lambeosaurid bone beds from the Upper Cretaceous Two Medicine Formation of Montana: taphonomic and biologic implications. Canadian Journal of Earth Sciences 30:997–1006.
Horner, J. R. 1994. "Steak knives, beady eyes, and tiny little arms (A portrait of T-rex as a scavenger)"; pp. 157–164 in G. D. Rosenberg and D. L. Wolberg (eds.), Dino-Fest. The Paleontological Society, Special Bulletin 7.
Horner, J. R. 1994. "Comparative taphonomy of some dinosaur and extant bird colonial nesting grounds" – Chapter 8; pp. 117–121 in K. Carpenter, C. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies. Cambridge University Press, New York, New York.
Horner, J. R. and Currie, P. J. 1994. "Embryonic and neonatal morphology and ontogeny of a new species of Hypacrosaurus (Ornithischia: Lambeosauridae) from Montana and Alberta"; pp. 312–336 in K. Carpenter, C. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies. Cambridge University Press, New York, New York.
Maxwell, D. and Horner, J. R. 1994. Neonate dinosaurian remains and dinosaurian eggshell from the Cloverly Formation, Montana. Journal of Vertebrate Paleontology 14(1):143–146.
Weishampel, D. B. and Horner, J. R. 1994. "Life history syndromes, heterochrony, and the evolution of Dinosauria"; pp. 229–243 in K. Carpenter, C. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies. Cambridge University Press, New York, New York.
Padian, K., Ricqlès, A. de, and Horner, J. R. 1995. Bone histology determines identification of a new fossil taxon of pterosaur (Reptilia: Archosauria). Compte Rendue Academie Science Paris 320(2):77–84.
Horner, J. R. and Weishampel, D. B. 1996. Correction to: A comparative embryological study of two ornithischian dinosaurs (1988). Nature 383:103.
Ruben, J. A., Hillenius, W. J., Geist, N. R., Leitch, A., Jones, T. D., Currie, P. J., Horner, J. R. , and Espe III, G. 1996. The metabolic status of some Late Cretaceous dinosaurs. Science 273:1204–1207.
Horner, J. R. 1997. Rare preservation of an incompletely ossified fossil embryo. Journal of Vertebrate Paleontology 17(2):431–434.
Schweitzer, M. H., Johnson, C, Zocco, G., Horner, J. R. , and Starkey, J. 1997. Preservation of biomolecules in cancellous bone of Tyrannosaurus rex. Journal of Vertebrate Paleontology 17(2):349–359.
Schweitzer, M. H., Marshal, M., Carron, K., Bohle, D. S., Busse, S. C., Arnaold, E. V., Barnard, D., Horner, J. R., and Starkey, J. R. 1997. Heme compounds in dinosaur trabecular bone. Proceedings of the National Academy of Sciences 94:6291–6296.
Varricchio, D. J., Jackson, F., Borkowski,J., and Horner, J. R. 1997. Nest and egg clutches of the dinosaur Troodon formosus and the evolution of avian reproductive traits. Nature 385:247–249.
Horner, J. R. and Clouse, V. 1998. "An undisturbed clutch of hadrosaur eggs from the Judith River Formation of Montana"; pp. 22–25 in 1st International Meeting on Dinosaur Paleobiology, Museu Nacional de Historia Natural, Universidade de Lisboa, Spain.
Horner, J. R., Ricqlès, A. de, and Padian, K. 1999. Variation in dinosaur skeletochronology indicators: implications for age assessment and physiology. Paleobiology 25(4):295–304.
Horner, J. R. 1999. Egg clutches and embryos of two hadrosaurian dinosaurs. Journal of Vertebrate Paleontology 19(4):607–611.
Schweitzer, M. H. and Horner, J. R. 1999. Intravascular microstructures in trabecular bone of Tyrannosaurus rex. Ann. Paléontol. 85(3):179–192.
Horner, J. R. 2000. Dinosaur reproduction and parenting. Annual Reviews of Earth and Planetary Sciences 28:19–45.
Horner, J. R., Ricqlès, A. de, and Padian, K. 2000. Long bone histology of the hadrosaurid Maiasaura peeblesorum: growth dynamics and physiology based on an ontogenetic series of skeletal elements. Journal of Vertebrate Paleontology 20(1):109–123.
Ricqlès, A. de, Padian, K., Horner, J. R., and Francillon-Vieillot, H. 2000. Paleohistology of the bones of pterosaurs (Reptilia: Archosauria): anatomy, ontogeny, and biomechanical implications. Zoological Journal of the Linnean Society 129:349–385.
Horner, J. R., Padian, K., and de Ricqlès, A. 2001. Comparative osteohistology of some embryonic and perinatal archosaurs: developmental and behavioral implications for dinosaurs. Paleobiology 27(1):39–58.
Horner, J. R. 2001. "Dinosaur Ethology"; pp. 416–419 in Briggs and Crowther (eds.), Palaeobiology II. Blackwell Science, Oxford, England.
Horner, J. R. 2001. "Dinosaur Behaviour"; A3318 in Encyclopedia of Life Sciences. Macmillan References Ltd.
Horner, J. R., Schmitt, J., Jackson, F., and Hannah, R. 2001. "Bones and rocks of the Two Medicine/Judith River clastic wedge complex of Montana"; pp. 3–13 in C.L. Hill (ed.), SVP Field Guidebook. Museum of the Rockies Occasional Paper #3, Bozeman, Montana.
Padian, K., A. J. de Ricqlès, and J. R. Horner. 2001. Dinosaurian growth rates and bird origins. Nature 412:405–408.
Rayfield, E. J., Norman, D. B., Horner, C. C., Horner, J. R., Smith, P. M. Thompson, J. and Upchurch, P. 2001. Cranial design and function in a large theropod dinosaur. Nature 409:1033–1037.
Ricqlès, A. de, Padian, K., and Horner, J. R. 2001. "The bone histology of basal birds in phylogenetic and ontogenetic perspectives"; pp. 411–426 in Jacques Gauthier and Lawrence F. Gall (eds.), New Perspectives on the Origin and Early Evolution of Birds. Peabody Museum of Natural History, Yale University, New Haven, Connecticut.
Horner, J.R. 2002. "Evidence of dinosaur social behavior" - Chapter 9; pp. 71–78 in J. G. Scotchmoor, B. B. Breithaupt, D. A. Springer & T. R. Fiorillo (eds.), Dinosaurs, The Science behind the Stories. American Geological Institute, Alexandria, Virginia.
Padian, K. and Horner, J. R. 2002. Typology versus transformation in the origin of birds. Trends in Ecology and Evolution 17(3):120–124.
Varricchio, D. J., Horner, J. R. and Jackson, F. 2002. Embryos and eggs for the Cretaceous theropod Troodon formosus. Journal of Vertebrate Paleontology 22(3):564–576.
Chin, K., Eberth, D.A., Schweitzer, M.H., Rando, T.A., Sloboda, W.J., and Horner, J.R. 2003. Remarkable preservation of muscle tissue within a tyrannosaurid coprolite from Alberta, Canada. Palaios 18:286–294.
Ricqlès, A. de, Padian, K. and Horner, J. R. 2003. On the bone histology of some Triassic pseudosuchian archosaurs and related taxa. Annales de Palèontologie 89:67–101.
Ricqlès, A. de, Padian, K., Horner, J. R. , Lamm, E.-T., and Myhrvold, N. 2003. Osteohistology of Confuciousornis sanctus (Theropoda: Aves). Journal of Vertebrate Paleontology 23(2):373–386.
Goodwin, M. and Horner, J. R. 2004. Cranial histology of pachycephalosaurs (Ornithischia: Marginocephalia) reveals transitory structures inconsistent with head-butting behavior. Paleobiology 30(2):253–267.
Horner, J. R. and Padian, K. 2004. Age and growth dynamics of Tyrannosaurus rex. Proceedings Royal Society (B) 271(1551):1875–1880.
Horner, J. R., Weishampel, D. B., and Forster, C. 2004. "Hadrosauridae"; pp. 438–463 in Weishampel, Dodson and Osmolska (eds.), The Dinosauria. University of California Press, Berkeley, California.
Padian, K. and Horner, J.R. 2004. "Dinosaur physiology"; pp. 660–671 in Weishampel, Dodson and Osmolska (eds.), The Dinosauria. University of California Press, Berkeley, California.
Padian, K., Horner, J. R. and de Ricqlés, A. 2004. Little dinosaurs and pterosaurs; the evolution of archosaurian growth strategies. Journal of Vertebrate Paleontology 24:555–571.
Channing, A. C., Schweitzer, M. H., Horner, J. R., and McEneaney, T. 2005. A silicified bird from Quaternary hot spring deposits. Proceedings of the Royal Society (B) 272:905–911.
Main, R. P, Ricqlès, A. de, Horner, J. R., and Padian, K. 2005. The evolution and function of thyreophoran dinosaur scutes: implications for plate function in stegosaurs. Paleobiology 31(2):291–314.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2005. Gender-specific reproductive tissue in ratites and Tyrannosaurus rex. Science 308:1456–1460.
Schweitzer, M. H., Wittmeyer, J. L., Horner, J. R., and Toporski, J.B. 2005. Soft-tissue vessels and cellular preservation in Tyrannosaurus rex. Science 307:1952–1955.
Goodwin, M., Clemmens, B., Horner, J. R., and Padian, K. 2006. The smallest known Triceratops skull: new observations on ceratopsid cranial anatomy and ontogeny. Journal of Vertebrate Paleontology 26(1):103–112.
Horner, J. R. and Goodwin, M. 2006. Major cranial changes in Triceratops. Proceedings of the Royal Society (B) 273:2757–2761.
Prieto-Marquez, A., Weishampel, D. B., and Horner, J. R. 2006. The dinosaur Hadrosaurus foulkii, from the Campanian of the East Coast of North America, with a reevaluation of the genus. Acta Palaeontol. Pol. 51(1):77–98.
Ricqlès, A. de, Horner, J. R., and Padian, K. 2006. The interpretation of dinosaur growth patterns. Trends in Ecology and Evolution 21(11):596–597.
Chinnery, B. and Horner, J. R. 2007. A new neoceratopsian dinosaur linking North American and Asian taxa. Journal of Vertebrate Paleontology 27(3):625–641.
Lipkin, C., Sereno, P. C., and Horner, J. R. 2007. The furcula in Suchomimus tenerensis and Tyrannosaurus rex (Dinosauria: Theropoda: Tetanurae). Journal of Paleontology 81(6):1523–1527.
Schweitzer, M. H., Elsey, R. M., Dacke, C. G., Horner, J. R., and Lamm, E.-T. 2007. Do egg-laying crocodilian (Alligator mississippiensis) archosaurs form medullary bone? Bone 40:1152–1158.
Schweitzer, M. H., Suo, Z., Avci, R., Asara, J. M., Allen, M. A. Arce, F. T., and Horner, J. R. 2007. Analysis of soft tissue from Tyrannosaurus rex suggest the presence of protein. Science 316:277–280.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2007. Soft tissue and cellular preservation in vertebrate skeletal elements from the Cretaceous to the present. Proceedings of the Royal Society (B) 274:183–197.
Cooper, L. N., Lee, A. H., Taper, M. L., and Horner, J. R. 2008. Relative growth rates of predator and prey dinosaurs reflect effects of predation. Proc. Royal Soc. B 275:2609–2615.
Horner, J. R. and Goodwin, M. 2008. Ontogeny of cranial epi-ossifications in Triceratops. Journal of Vertebrate Paleontology 28(1):134–144.
Ricqlès, A. de, Padian, K., Knoll, F., and Horner, J. R., 2008. On the origin of high growth rates in archosaurs and their ancient relatives: Complementary histological studies on Triassic archosauriformes and the problem of a phylogenetic signal in bone histology. Annales de Paléontologie 94:57–76.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2008. "One pretty amazing T. rex." in Larson and Carpenter (eds.), Tyrannosaurus rex, The Tyrant King. Indiana University Press, Bloomington, Indiana.
Horner, J. R. and Goodwin, M. B. 2009. Extreme cranial ontogeny in the Upper Cretaceous dinosaur Pachycephalosaurus. PLoS ONE 4(10):e7626
Horner, J. R., Ricqlès, A. de, Padian, K., and Scheetz, R. 2009. Comparative long bone histology of the “hypsilophodontid” dinosaurs Orodromeus makelai, Dryosaurus altus, and Tenontosaurus tillettii (Ornithischia: Euornithopoda). Journal of Vertebrate Paleontology 29(3):1–14.
Schweitzer, M. H., Zheng, W., Organ, C. L., Avci, R., Suo, Z., Freimark, L. M., Lebleu, V. S., Duncan, M. B., Vander Heiden, M. G., Neveu, J. M., Lane, W. S., Cottrell, J. S., Horner, J. R., Cantly, L. C., Kalluri, R., and Asara, J. M. 2009. Biomolecular characterization and protein sequences of the Campanian hadrosaurs B. Canadensis. Science 324:626–631.
Wolff, E. D. S., Salisbury, S. W., Horner, J. R., and Varricchio, D. J. 2009. Common avian infection plagued the tyrant dinosaurs. PLoS ONE 4(9):e7288.
Archibald, J. D., et al. (29 authors). 2010. Cretaceous extinctions: Multiple causes. Science 328:973
Goodwin, M. B. and Horner, J. R. 2010. "Historical collecting bias and the fossil record of Triceratops in Montana"; pp. 551–563 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs. The Royal Tyrrell Museum Ceratopsian Symposium, Indiana University Press, Bloomington, Indiana.
Jackson, F. D., Horner, J. R., and Varricchio, D. J. 2010. A study of a Troodon egg containing embryonic remain using epifluorescence microscopy and other techniques. Cretaceous Research 31:255–262.
Longrich, N. R., Horner, J. R., Erickson, G. M., and Currie, P. J. 2010. Cannibalism in Tyrannosaurus rex. PLoS ONE 5(10):e13419.
McDonald, A. T. and Horner, J. R. 2010. "New material of “Styracosaurus” ovatus from the Two Medicine Formation of Montana." pp. 156–168 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs. The Royal Tyrrell Museum Ceratopsian Symposium, Indiana University Press, Bloomington, Indiana.
Scannella, J. B. and Horner, J. R. 2010. Torosaurus Marsh, 1891, is Triceratops Marsh, 1889 (Certatopisdae: Chasmosaurinae): synonomy through ontogeny. Journal of Vertebrate Paleontology 30(4):1157–1168.
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Therrien, F., and Horner, J. R. 2011. A Subadult skull of Hypacrosaurus stebingeri (Ornithischia: Lambeosaurinae): anatomy and comparison. Historical Biology 23(1):63–72.
Horner, J. R., Goodwin, M., and Myhrvold, N. 2011. Dinosaur census reveals abundant Tyrannosaurus and rare ontogenetic stages in the Upper Cretaceous Hell Creek Formation (Maastrichtian), Montana, USA. PLoS ONE 6(2):e16574.
Horner, J. R. and Lamm, E-T. 2011. Ontogeny of the parietal frill of Triceratops: a preliminary histological analysis. Comptes Rendus Palevol 10:439–452.
Fowler, D. W., Woodward, H. N., Freedman, E. A., Larson, P. L., and Horner, J. R. 2011. Reanalysis of “Raptorex kriegsteini”: a juvenile tyrannosaurid dinosaur from Mongolia. PLoS ONE 6(6):e21376.
Gates, T. A., Horner, J. R., Hanna, R. R., and Nelson, C. R. 2011. New unadorned hadrosaurine hadrosaurid (Dinosauria, Ornithopoda) from the Campanian of North America. Journal of Vertebrate Paleontology 31(4):798–811.
Padian, K. and Horner, J. R. 2011. The evolution of ‘Bizarre structures’ in dinosaurs: biomechanics, sexual selection, social selection or species recognition? Journal of Zoology 283(1):3–17. doi:10.1111/j.1469-7998.2010.00719.x
Padian, K. and Horner, J. R. 2011. The definition of sexual selection and its implications for dinosaurian biology. Journal of Zoology. 283(1):23–27. doi:10.1111/j.1469-7998.2010.00761.x
Prieto-Marquez, A., Minjin, B., and Horner, J. R. 2011. A diminutive deinonychosaur (Dinopsauria: Theropoda) from the Early Cretaceous of Oosh (Ovorkhangai, Mongolia). Alcheringa 36(1):117-136. doi:10.1080/03115518.2011.590401
Scannella, J. B. and Horner, J. R. 2011. ‘Nedoceratops’: An example of a transitional morphology. PLoS ONE 6(12):e28705. doi:10.1371/journal.pone.0028705
Schott, R. K., Evans, D. C., Goodwin, M. B. Horner, J. R., Brown, C. M., and Longrich, N. R. 2011. Cranial ontogeny in Stegoceras validum (Dinosauria: Pachycephalosauria): a quantitative model of pachycephalosaur dome growth and variation. PLoS ONE 6(6):e21092. doi:10.1371/journal.pone.0021092
Woodward, H. N., Horner, J. R., and Farlow, J. O. 2011. Osteohistological evidence for determinate growth in the American Alligator. Journal of Herpetology 45(3):339–342.
Bailleul, A. M., Hall, B. K., and Horner, J. R. 2012. First evidence of dinosaurian secondary cartilage in the post-hatching skull of Hypacrosaurus stebingeri (Dinosauria, Ornithischia). PLoS ONE 7(4):e36112. doi:10.1371/journal.pone.0036112
Horner, J. R. 2012. Dinosaur behaviour. in ELS. John Wiley & Sons, Ltd: Chickester. doi:10.1002/9780470015902.a0003318.pub3
Zelenitsky, D. K., Horner, J. R., and Therrien, F. 2012. Dinosaur Eggs: pp. 613–620 in M. K. Brett-Surman, T. R. Holtz and J.O. Farlow (eds.), The Complete Dinosaur. Indiana University Press, Bloomington, Indiana.
Padian, K. and Horner, J. R. 2013. The species recognition hypothesis explains exaggerated structures in non-avialan dinosaurs better than sexual selection does. Comptes Rendus Palevol 13(2): doi:10.1016/j.crpv.2013.10.004
Bailleul, A. M., Hall, B. K., Horner, J. R. 2013. Secondary cartilage revealed in a non-avian dinosaur embryo. PLoS ONE 8(2):e56937. doi:10.1371/journal.pone.0056937
Padian, K. and Horner, J. R. 2013. Misconceptions of sexual selection and species recognition: a response to Knell et al. and to Mendelson and Shaw. Trends in Ecology & Evolution 28(5): doi:10.1016/j.tree.2013.01.011
Goodwin, M. B. and Horner, J. R. 2014. Cranial morphology of a juvenile Triceratops skull from the Hell Creek Formation, McCone County, Montana, with comments on the fossil record of ontogenetically younger skulls. Special Paper of the Geological Society of America 503:333-347. doi:10.1130/2014.2503(13)
Horner, J. R. 2014. Paleontology: a cock's comb on a duck-billed dinosaur. Current Biology 24(2):R85-6. doi:10.1016/j.cub.2013.12.007
Woodward, H. N., Horner, J. R., Farlow, J. O. 2014. Quantification of intraskeletal histovariability in Alligator mississippiensis and implication for vertebrate osteohistology. PeerJ 2:e422. doi:10.7717/peerj.422
Scannella, J. B., Fowler, D. W., Goodwin, M. B., Horner, J. R. 2014. Evolutionary trends in Triceratops from the Hell Creek Formation, Montana. Proceedings of the National Academy of Sciences 111(28):10245-10250. doi:10.1073/pnas.1313334111
Rashid, D. J., Chapman, S. C., Larsson, H. C. E., Organ, C. L., Bebin, A. G., Merzdorf, C. S., Bradley, R., Horner, J. R. 2014. From dinosaurs to birds: a tail of evolution. EvoDevo 5(25):1-20. doi:10.1186/2041-9139-5-25
Padian, K. and Horner, J. R. 2014. Darwin's sexual selection: understanding his ideas in context. Comptes Rendus Palevol 13(8). doi:10.1016/j.crpv.2014.09.001
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Horner, J. R., Therrien, F. 2014. Cranial morphology and variation in Hypacrosaurus stebingeri (Ornithischia: Hadrosauridae): chapter 14: pages 245-276 in Eberth, D. A. and Evans, D. C (eds.). Hadrosaurs. Indiana University Press, Bloomington, Indiana.
Bailleul, A. M., Nyssen-Behets, C., Lengele, B., Hall, B. K., Horner, J. R. 2015. Chondroid bone in dinosaur embryos and nestlings (Ornithischia: Hadrosauridae): insights into the growth of the skull and the evolution of skeletal tissues. Comptes Rendus Palevol (in press). doi:10.1016/j.crpv.2014.12.004
Horner, J. R., Woodward, H. N., Bailleul, A. M. 2015. Mineralized tissues in dinosaurs interpreted as having formed through metaplasia: a preliminary evaluation. Comptes Rendus Palevol (in press). doi:10.1016/j.crpv.2015.01.006
Padian, K., Werning, S., Horner, J. R. 2015. A hypothesis of differential secondary bone formation in dinosaurs. Comptes Rendus Palevol
(in press). doi:10.1016/j.crpv.2015.03.002
Cubo, J., Woodward, H. N., Wolff, E., Horner, J. R. 2015. First reported cases of biomechanically adaptive bone modeling in non-avian dinosaurs. PLoS ONE 10(7). doi:10.1371/journal.pone.0131131
Published Abstracts (102):
Baird, D. and Horner, J. R. 1977. A fresh look at the dinosaurs of New Jersey and Delaware. New Jersey Acad. Sci. Bulletin 22(2):50.
Lund, R. and Horner, J. R. 1979. Biotic distribution and diversity in the Bear Gulch Basin. Ninth International Congress of Carboniferous Stratigraphy and Geology Abstracts:120.
Melton, W. G., Lund, R. and Horner, J. R. 1979. Faunal distribution within the Bear Gulch lentil. Ninth International Congress of Carboniferous Stratigraphy and Geology Abstracts:136.
Horner, J. R. 1987. Evolution of terrestrial Late Cretaceous faunal communities. Journal of Vertebrate Paleontology 7(3, Supplement):18A.
Currie, P. J. and Horner, J. R. 1988. Lambeosaurine hadrosaur embryos (Reptilia: Ornithischia). Journal of Vertebrate Paleontology 8 (3, Supplement):13A.
Horner, J. R. 1988a. Morphological evidence of mammalian-grade herd structure in hadrosaurian and ceratopsian dinosaurs. International Symposium on Vertebrate Behavior as Derived from the Fossil Record 1988:13. Montana State University, Bozeman, Montana.
Horner, J. R. 1989b. The ontogenetic predictability of hadrosaur crania. Journal of Vertebrate Paleontology 9(3, Supplement):25A.
Horner, J. R. 1988c. Cranial allometry of Maiasaura peeblesorum (Ornithischia; Hadrosauridae) and its behavioral significance. Journal of Vertebrate Paleontology 8(3, Supplement):18A.
Cooley, J., Schmitt, J., and Horner, J. R. 1992. Fluvial systems of the Upper Jurassic Morrison Formation, Gallatin and Beartooth Ranges, SW Montana. SEPM Annual Meeting Bulletin 1992.
Horner, J. R. 1992b. Dinosaur behavior and growth. Paleontological Society Special Publication 6:135.
Barreto, C. B., Albrecht, R. M., Horner, J. R., and Wilsman, N. H. 1993. Evidence of dinosaur growth plate morphology in the fossil record. The Society for Orthopedic Research, Annual Meeting 1993.
Barreto, C. B., Albrecht, R. M., Horner, J. R., and Wilsman, N. H. 1993. Dinosaur long bone growth plates. Journal of Vertebrate Paleontology 13(3, Supplement):25A.
Clouse, V. and Horner, J. R. 1993. Eggs and embryos from the Judith River Formation of Montana. Journal of Vertebrate Paleontology 13(3, Supplement):42A.
Horner, J. R. 1993. Ontogenetic development of the lambeosaurid narial crest. Journal of Vertebrate Paleontology 13(3, Supplement):42A.
Padian, K., Horner, J. R., and de Ricqlès, A. 1993. A new azhdarchid pterosaur from the Two Medicine Formation (Late Cretaceous: Campanian) of Montana. Journal of Vertebrate Paleontology 13(3, Supplement):52A
Ricqlès, A. de, Padian, K., and Horner, J. R. 1993. Paleohistology of pterosaur bones. Journal of Vertebrate Paleontology 13(3, Supplement):54A.
Schweitzer, M. H., Cano, R. J., Starkey, J. R., and Horner, J. R. 1994. Multiple lines of evidence for the preservation of collagen and other biomolecules in unmineralized bone from Tyrannosaurus rex. Journal of Vertebrate Paleontology 14(3, Supplement):45A.
Carter, D. R., Mikic, B., and Horner, J. R. 1995.Growth plate differences between mammals and dinosaurs are related to both genotypic and mechanical factors. Journal of Vertebrate Paleontology 15(3, Supplement):22A
Horner, J. R. 1995. Morphology and function of the enclosed narial chambers of lambeosaurid dinosaurs. Journal of Vertebrate Paleontology 15(3, Supplement):36A.
Horner, J. R. 1996a. Transgressions, regressions, and the evolution of dinosaurs. The Paleontological Society, Special Publication 8:178.
Horner, J. R. 1996b. New evidence for post-eclosion parental attention in Maiasaura peeblesorum. Journal of Vertebrate Paleontology 16(3, Supplement):42A.
Laws, R., Hasiotis, S. T., Fiorillo, A. R., Chure, D., Breithaupt, B. H., and Horner, J. R. 1996. The demise of a Jurassic Morrison dinosaur after death - three cheers for the dermestid beetle. Geological Society of America, Abstracts with Programs 28:530A.
Goodwin, M. B., Clemens, W. A., and Horner, J. R. 1997. Morphological variation and ontogeny in the skull of Triceratops. Journal of Vertebrate Paleontology 17(3, Supplement):49A.
Horner, J. R., Ricqlès, A. de, and Padian, K. 1997. Histological analysis of a dinosaur skeleton: evidence of skeletal growth variation. Journal of Morphology 232(3):267.
Horner, J. R. and Varricchio, D. J. 1997. Embryonic remains of Troodon formosus from the Two Medicine Formation of Montana. Journal of Vertebrate Paleontology 17(3, Supplement):53A.
Maxwell, W., Hallas, B. H., and Horner, J. R. 1997. Further neonate dinosaurian remains and dinosaurian eggshell from the Cloverly Formation of Montana. Journal of Vertebrate Paleontology 17(3):63A.
Padian, K., Ricqlès, A. de, and Horner, J. R. 1997. Pterosaur bone histology: growth rates and mechanics. Journal of Morphology 232(3):306.
Ricqlès, A. de, Horner, J. R., and Padian, K. 1997. Histological evidence of dinosaur growth patterns. Abstracts, Third World Herpetological Congress, Prague.
Ricqlès, A. de, Padian, K., and Horner, J. R. 1997. Comparative biology and the bone histology of extinct tetrapods: what does it tell us? Journal of Morphology 232(3):246.
Varricchio, D. J., Jackson, F., Borkowski, J. J., and Horner, J. R. 1997. Coelurosaurian eggs: between a rock and a soft place. Journal of Vertebrate Paleontology 17(3, Supplement):82A.
Ancell, C. A., Harmon, R., and Horner, J. R. 1998. Gar in a duckbill: preservation of a scavenger and its prey. Journal of Vertebrate Paleontology 18(3, Supplement):24A
Cooper, L. N., Lamm, E., Horner, C. C. and Horner, J. R. 1998. Annual changes in mass of Hypacrosaurus stebingeri as hypothesized from lines of arrested growth. Journal of Vertebrate Paleontology 18(3, Supplement):39A
Horner, J. R. and Goodwin, M. B. 1998. Did pachycephalosaurs really head-butt? An osteohistological cranial analysis. Journal of Vertebrate Paleontology 18(3, Supplement):52A
Ricqlès, A. de, Horner, J. R., and Padian, K. 1998. Growth dynamics of the hadrosaurid dinosaur Maiasaura peeblesorum. Journal of Vertebrate Paleontology 18(3, Supplement):72A.
Schmitt, J. G., Horner, J. R., Laws, R. R. and Jackson, F. 1998. Debris-flow deposition of a hadrosaur-bearing bone-bed, Upper Cretaceous Two Medicine Formation, northwest Montana. Journal of Vertebrate Paleontology 18(3, Supplement):76A.
Cooper, L. N. and Horner, J. R. 1999. Growth rate of Hypacrosaurus stebingeri as hypothesized from lines of arrested growth and whole femur circumference. Journal of Vertebrate Paleontology 19(3, Supplement):39A.
Hanna, R. R., LaRock, J. W., and Horner, J. R. 1999. Pathological brachylophosaur bones from the Upper Cretaceous Judith River Formation, northeastern Montana. Journal of Vertebrate Paleontology 19(3, Supplement):49A.
Horner, J. R. 1999. Dinosaur biozones in the Campanian clastic wedge complex of Montana, and their biological implications. AAPG, Rocky Mountain Section, Abstracts with Program 1999:A6.
Horner, J. R., Padian, K., and de Ricqlès, A. 1999. Osteohistology of some embryonic and perinatal archosaurs: phylogenetic and behavioral implications for dinosaurs. Journal of Vertebrate Paleontology 19(3, Supplement):53A.
Padian, K., Horner, J. R., and de Ricqlès, A. 1999. Dinosaurian growth rates and the evolution of life history strategies. Journal of Vertebrate Paleontology 19(3, Supplement):67A.
Ricqlès, A. de, Padian, K., and Horner, J. R. 1999. The bone histology of basal birds in phylogenetic and ontogenetic perspectives. Journal of Vertebrate Paleontology 19(3, Supplement):70A.
Main, R. P., Padian, K, and Horner, J. R. 2000. Comparative histology, growth, and evolution of archosaurian osteoderms: why did Stegosaurus have large dorsal plates? Journal of Vertebrate Paleontology 20(3):56A.
Barreto, C., Marshall, C., and Horner, J. R. 2001. Differential and determinate growth in dinosaurs and other reptiles. Journal of Vertebrate Paleontology 21(3):31A.
Goodwin, M. and Horner, J. R. 2001. How Triceratops got its horns: New information from a growth series on cranial morphology and ontogeny. Journal of Vertebrate Paleontology 21(3):56A.
Higby Schweitzer, M., Schmitt, J.G., 2001, and Horner, J.R. 2001. Paleoenvironmental and taphonomic parameters of an exceptionally preserved Tyrannosaurus rex. Geological Society of America Abstracts 33(6):388.
Horner, C. C., Horner, J. R., and Weishampel, D. B. 2001.Comparative internal cranial morphology of some hadrosaurian dinosaurs using computerized tomographic X-ray analysis and rapid prototyping. Journal of Vertebrate Paleontology 21(3):64A
Horner, J. R. and Padian, K. 2001. Evolution of archosaurian growth strategies. Journal of Vertebrate Paleontology 21(3):63A
Jerison, H. J., Horner, J. R., and Horner, C. C. 2001. Dinosaur forebrains. Journal of Vertebrate Paleontology 21(3):65A.
Padian, K., Horner, J. R., de Ricqlès. 2001. Dinosaurian life history strategies: evidence of differential growth rates from bone histology. American Zoologist 40(5).
Ricqlès, A de, Padian, K., and Horner, J. R. 2001. Bone histology in extant and extinct archosauromorphs: A structurofunctional exploration in ontogeny and phylogeny. Journal of Morphology 248(3):222.
Horner, J. R. and Marshall, C. 2002. Keratin covered dinosaur skulls. Journal of Vertebrate Paleontology 22(3):67A.
Padian, K, Horner, J. R., and Ricqlès, A. 2002. How dinosaurs grew: growth rates and strategies. Geological Society of America.
Ricqlès, A. de, Padian, K., and Horner, J. R. 2002. On the bone histology of some Triassic pseudosuchian archosaurs and related taxa: a preliminary report. 8th International Symposium on Mesozoic Terrestrial Ecosystems, Cape Town, South Africa.
Chinnery, B. and Horner, J. R. 2003. New basal neoceratopsioan from the lower Two Medicine Formation of Montana provides a link between Asian and North America taxa. Journal of Vertebrate Paleontology 23(3):40A.
Prieto-Marquez, A., Weishampel, D. B., and Horner, J. R. 2003. Taxonomy and systematics of the holotype of Hadrosaurus foulkii (Dinosauria, Ornithopoda) from the Late Cretaceous of eastern North America. Journal of Vertebrate Paleontology 23(3):88A.
Schweitzer, M., Channing, A., Horner, J. R., and Chiappe, L. 2003. Microbial biomineralization and exceptional preservation of vertebrate fossils. Journal of Vertebrate Paleontology 23(3):95A.
Wolff, E. and Horner, J. R. 2003. The first occurrence of Orodromeus makelai postcrania from the Late Campanian Judith River Formation of eastern Montana. Journal of Vertebrate Paleontology 23(3):112A.
Horner, J. R. and Padian, K. 2004. Age and growth dynamics of Tyrannosaurus rex. Journal of Vertebrate Paleontology 24 (3):72A.
Padian, K., Horner, J. R., and Dhaliwal, J. 2004. Species recognition as the principal cause of bizarre structures in dinosaurs. Journal of Vertebrate Paleontology 23(3):100A.
Schweitzer, M., Wittmeyer, J., and Horner, J. R. 2004. A novel dinosaurian tissue exhibiting unusual preservation. Journal of Vertebrate Paleontology 24(3):111A.
Currie, P. J., Henderson, M., Horner, J. R., and Williams, S. A. 2005. On tyrannosaur teeth, tooth positions and the taxonomic status of Nanotyrannus lancensis. The Origin, Systematics, and Paleobiology of Tyrannosauridae Symposium Abstracts:19. Burpee Museum, Rockford, Illinois
Horner, J. R. and Goodwin, M. 2005. A new Triceratops cranial growth series. Journal of Vertebrate Paleontology 25(3):71A.
Padian, K., Horner, J. R., and Lee, A. 2005. Sexual dimorphism in dinosaurs? A review of the evidence. Journal of Vertebrate Paleontology 25(3):98A.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2005. Reproductive tissues in exceptionally well preserved Tyrannosaurus rex (MOR 1125). The Origin, Systematics, and Paleobiology of Tyrannosauridae Symposium Abstracts:30, Burpee Museum, Rockford, Illinois.
Horner, J. R. 2006. Osteohistology in dinosaur research. AAAS symposium, St. Louis, MO.
Padian, K., de Ricqlès, A, and Horner, J. R. 2006. Comparative long bone histology and growth of some hypsilophodontid dinosaurs. Journal of Vertebrate Paleontology 26(3):108A.
Reizer, J, and Horner, J. R. 2006. An ontogenetic series of the ceratopsid dinosaur Einiosaurus procurvicornis as determined by long bone histology. Journal of Vertebrate Paleontology 26(3):114A.
Ricqlès, A. de, Padian, K., and Horner, J. R. 2006. Complementary studies on archosauriform and basal archosaur bone histology: is research on a plesiomorphic histological condition warranted? Chinese Symposium.
Zelenitsky, D., Therrien, F., Currie, P., Henderson, D., and Horner, J. R. 2006. Locomotory behavior in the lambeosaurine Hypacrosaurus stebingeri. Journal of Vertebrate Paleontology 26(3):143A.
DeBoef, M., Larsson, H., and Horner, J. R. 2007. Measurements of vasculature in Tyrannosaurus rex show a relationship between growth rate and vascular orientation. Journal of Vertebrate Paleontology 27(3):76A.
Horner, J. R., Goodwin, M. B., and Woodward, H. 2007. Synonymy consequences of dinosaur cranial ontogeny. Journal of Vertebrate Paleontology 27(3):92A.
Brink, K., Zelenitsky, D., Evans, D., and Horner, J.R. 2008. New subadult Hypacrosaurus stebingeri skulls and a comparative analysis of allometric crest growth in lambeosaurines. Journal of Vertebrate Paleontology 28(3):56A.
Evans, D., Witmer, L., Ridgely, R., and Horner, J. R. 2008. Endocranial anatomy of lambeosaurine dinosaurs: implications for cranial crest function and evolution. Journal of Vertebrate Paleontology 28(3):75A.
Lee, A., Cooper, L., Taper, M., and Horner, J. R. 2008. Rapid growth of the hadrosaur Hypacrosaurus reflects direct and indirect effects of predation. Journal of Vertebrate Paleontology 28(3):104A.
Horner, J. R. 2008. Dinosaur Cranial Ontogeny. Paleofest, Burpee Museum, Rockford, Illinois.
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Horner, J. R., and Therrien, F. 2009. Ontogenetic development of the skull of Hypacrosaurus stebingeri. Fourth International Symposium on Dinosaur Eggs and Babies. Museum of the Rockies Occasional Paper #4:12, Bozeman, Montana.
Goodwin, M. B. and Horner, J. R. 2009. The fossil record of Triceratops from the Hell Creek Formation, Montana. pp. 147 in 9th North American Paleontological Convention, Abstracts. Cincinnati Museum Center Scientific Contributions #3, Cincinnati, Ohio.
Horner, J. R., Goodwin, M. B. and Myhrvold, N. 2009. Hell Creek Formation dinosaur census reveals abundant Tyrannosaurus. pp. 146 in 9th North American Paleontological Convention, Abstracts. Cincinnati Museum Center Scientific Contributions #3, Cincinnati, Ohio.
Horner, J. R. and Lamm, E-T. 2009. Metaplasia provided dinosaur skulls extreme morphological plasticity during ontogeny. Journal of Vertebrate Paleontology 29(3):117A.
Jackson, F. D., Horner, J. R., and Varricchio, D. 2009. A study of a Troodon egg containing embryonic remains using epiflourescence microscopy and other techniques. Fourth International Symposium on Dinosaur Eggs and Babies. Museum of the Rockies Occasional Paper #4:20, Bozeman, Montana.
Jackson, F. D., Horner, J. R., and Varricchio, D. 2009. A study of a Troodon egg containing embryonic remains using epiflourescence microscopy and other techniques. Journal of Vertebrate Paleontology 29(3):121A.
Padian, K., Horner, J. R., Fowler, D., and Scannella, J. 2010. How a synergy of species recognition and social signaling explains cranial anatomy and ontogeny in several groups of dinosaurs. Journal of Vertebrate Paleontology 30(4, Supplement):143A.
Woodward, H. and Horner, J. R. 2010. Osteohistological analysis of Alligator mississippiensis indicates absence of fibrolamellar bone in crocodilians and confirms determinate growth with the first report of external fundamental system: implications for tetrapod histology. Journal of Vertebrate Paleontology 30(4, Supplement):190A.
Baillieu, A. and Horner, J. R. 2011. Early ossification and calcified tissues in the skull of Hypacrosaurus stebingeri (Ornithischia, Lambeosaurinae): a preliminary study. pp. 11 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology, Drumheller, Alberta, Canada.
Bailleul, A. M. and Horner, J. R. 2011. Perinatal histomorphogenesis of craniofacial elements of Hypacrosaurus stebingeri (Ornithischia, Lambeosauridae): a preliminary study. Vol.1:pp. 57 in International Symposium on Paleohistology, Abstracts (M. Kohler, X. Jordana and N. M. Moratalla Eds), Institut Català de Paleontologia, Barcelona, Spain.
Bailleul, A. and Horner, J. R. 2011. Early ossification and calcified tissues in the skull of Hypacrosaurus stebingeri (Ornithischia, Lambeosaurinae): a preliminary study. Journal of Vertebrate Paleontology 31(5, abstracts vol):65.
Baker, K., Scannella, J., Hall, L., and Horner, J. R. 2011. Biogeographic implications of a partial ceratopsid skeleton from the lower Two Medicine Formation (Campanian), Montana. Journal of Vertebrate Paleontology 31(5, abstracts vol):65.
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Horner, J. R., and Therrien, F. 2011. Hypacrosaurus stebingeri: an exceptional record of dinosaurian cranial ontogeny. pp. 14-15 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology.
Evans, D. C., Currie, P. J., Witmer, L. M., and Horner, J. R., and Ebreth, D. 2011. A new low crested lambeosaurine hadrosaurid from the Dinossaur Park Formation of Sandy Point, eastern Alberta. pp. 48–49 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology, Drumheller, Alberta, Canada.
Fowler, D., Scannella, J., and Horner, J. R. 2011 Reassessing ceratopsid diversity using unified frames of reference. Journal of Vertebrate Paleontology 31(5, abstracts vol):111.
Goodwin, M., Stanton, K., Horner, J. R., and Carlson, S. 2011. Oxygen isotopic variability and preservation in Tyrannosaurus rex, modern ratites and crocodilians: revisiting the thermophysiology of T.rex using δ180. Journal of Vertebrate Paleontology 31(5, abstracts vol):117.
Horner, J. R. and Woodward, H. N. 2011. Riddle of the humongous hadrosaurs: what these giants reveal about dinosaur ontogeny, evolution, and ecology. Vol.1:pp. 66 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology, Drumheller, Alberta, Canada.
Horner, J. R., Woodward, H. N., and Bailleul, A. 2011. Evaluating maturity in dinosaurs. Vol.1:pp. 53 in International Symposium on Paleohistology, Abstracts (M. Kohler, X. Jordana and N. M. Moratalla Eds), Institut Català de Paleontologia, Barcelona, Spain.
Lawver, D., Jackson, F., and Horner, J. R. 2011. Turtle eggs and embryos in a clutch from the Upper Cretaceous (Campanian) Judith River Formation of Montana. Journal of Vertebrate Paleontology 31(5, abstracts vol):141.
Scannella, J., Fowler, D., Trevethan, I., Roberts, D., and Horner, J. R. 2011. Individual variation in Triceratops from the Hell Creek Formation, Montana: implications for dinosaur taxonomy. Journal of Vertebrate Paleontology 31(5, abstracts vol):187.
Woodward, H. N., and Horner, J. R. 2011. A robust single element microanalysis of a hadrosaurid dinosaur (Ornithopoda: Maiasaura peeblesorum) population reveals ontogenetic trends in growth and development. Vol.1:pp. 56 in International Symposium on Paleohistology, Abstracts (M.Kohler, X. Jordana and N. M. Moratalla Eds), Institut Català de Paleontologia, Barcelona, Spain.
Fowler, D. W., Scannella, J. B., Goodwin, M. G., and Horner, J. R. 2012. How to eat a Triceratops: large sample of toothmarks provides new insight into the feeding behavior of Tyrannosaurus. Journal of Vertebrate Paleontology 32(5, abstracts vol):96.
Horner, V. and Horner, J. R. 2012. Sturgeon dorsal osteoderm ontogeny: A transformational model for marginocephalian squamosal ornaments. Journal of Vertebrate Paleontology 32(5, abstracts vol):113.
Scannella, J. B., Fowler, D.W., Goodwin, M. G., and Horner, J. R. 2012. Transitional Triceratops: Details of an ontogenetic sequence from the upper Middle Unit of the Hell Creek Formation, Montana. Journal of Vertebrate Paleontology 32(5, abstracts vol):166.
Woodruff, C., and Horner, J. R. 2012. A re-evaluation of the vertebral Soft Tissue reconstruction within Dinosauria based on alternate extant analogues. Journal of Vertebrate Paleontology 32(5, abstracts vol):195.
Woodward, H.N., Horner J. R., and Farlow, J.O. 2012. Paleobiological implications of the growth history and histovariability in a population of the hadrosaurid dinosaur Maiasaura peeblesorum. Journal of Vertebrate Paleontology 32(5, abstracts vol):196.
Boatman, E. M., Goodwin, M. B., Holman, H., Fakra, S., Schweitzer, M. H., Gronsky, R., Horner, J. R. 2014. Synchrotron chemical and structural analysis of Tyrannosaurus rex blood vessels: the contribution of collagen hypercrosslinking to tissue longevity. Microscopy and Microanalysis 20(Suppl 3):1430-1431.
Edited Book (1):
Carpenter, K., Hirsch, K. F., and Horner, J. R. (eds.). 1994. Dinosaur Eggs and Babies, Cambridge University Press, New York, 372 pp.
Popular Books (8):
Horner, J. R. and J. Gorman. 1985. Maia, a Dinosaur Grows Up. Museum of the Rockies, Bozeman, Montana & Running Press, Philadelphia Pensylvannia, 34 pp.
Horner, J. R. and J. Gorman. 1988. Digging Dinosaurs. Workman Publishing, New York, 210 pp.
Horner, J. R. and D. Lessom. 1992. Digging up Tyrannosaurus rex. Crown Publishers, New York, 36 pp.
Horner, J. R. and D. Lessom. 1993. The Complete T-rex. Simon and Schuster, New York, 235 pp.
Horner, J. R. and E. Dobb. 1997. Dinosaur Lives. Harper Collins, New York, 244 pp.
Horner, J. R. 2001. Dinosaurs Under the Big Sky. Mountain Press, Missoula, Montana, 194 pp.
Horner, J. R. 2007. Digging Up Dinosaurs. Farcountry Press, Helena, Montana, 49pp.
Horner, J. R. and J. Gorman. 2009. How to Build a Dinosaur; Extinction Doesn’t Have to be Forever. Dutton, New York, 246 pp.
Popular Publications (46-Total # Articles)
Horner, J. R. 1982. "Coming home to roost." Montana Outdoors 13(4):2–5.
Horner, J. R. 1984. "The nesting behavior of dinosaurs." Scientific American 250(4):130–137.
Horner, J. R. and D. B. Weishampel. 1989. "Dinosaur eggs: the inside story." Natural History 89(12):60–67.
Horner, J. R. 1996-2001 (bi-monthly column) Wild Outdoor World, Rocky Mountain Elk Foundation, Children’s magazine, Montana.
Horner, J. R. 1997. "Behavior"; pp. 45–50 in P. Currie and K. Padian (eds.), The Encyclopedia of Dinosaurs. Academic Press, San Diego, California.
Horner, J. R. 1997. "Willow Creek Anticline"; pp. 786 in P. Currie and K. Padian (eds.), The Encyclopedia of Dinosaurs. Academic Press, San Diego, California.
Horner, J. R. 1998. "Dinosaur behavior." National Forum 78(3):12–15.
Horner, J. R. 2000. "Dinosaur." Encyclopedia Americana 9:123–131.
Horner, J. R. 2000. "Hadrosaur." Encyclopedia Americana ###
Horner, J. R. 2004. Jack Horner: "An Intellectual Autobiography." in Montana Professor , 14(2):15–19. University of Montana, Missoula, Montana.
Horner, J. R., Padian, K, and de Ricqlès, A. 2005. "How dinosaurs grew so large and so small." Scientific American 293(4):32–39.
Horner, J. R. 2005. "Dinosaur Behaviour", Encyclopedia of Life Sciences, John Wiley & Sons, Ltd.
Horner, J. R. 2007. "Discovery of Maiasaura." Cosmos Magazine.
Horner, J. R. 2008. "The extraordinary characteristics of dyslexia." Perspectives on Language and Literacy 34:35. International Dyslexia Association.
Horner, J. R. 2009. "Dinosaurs at the table" - Chapter 6; pp. 111–121 in D. Seplowski and M. Ruse (eds.) The Paleobiological Revolution; Essays on the Growth of Modern Paleontology. University of Chicago Press, Chicago, Illinois.
Horner, J. R. and J. Gorman. 2009. "Dinosaur Resurrection." Discover Magazine, April 2009:50–53.
Horner, J. R. 2012. "Why do animals that fly have feathers, not including bats"?; pp. 99–100 in Gemma Elwin Harris (ed.), Big Questions from Little People. Farber & Farber Lmt., London.
Horner, J. R. and Horner, V. S. 2012. "Time traveling in Montana’s Mesozoic park." in Distinctly Montana, Fall, 2012.
Horner, J. R. and Druckenmiller, P. 2012. "Polar Dinosaurs"; pp. 127–130 in D. S. Miller (ed.), On Arctic Ground. Braid River Press, Seattle, Washington.
Schram, F. R. and Horner, J. R. 1978. Crustacea of the Mississippian Bear Gulch Limestone of central Montana. Journal of Paleontology 52(2):394–406.
Baird, D. and Horner, J. R. 1979. Cretaceous dinosaurs of North Carolina. Brimleyana 2:1–28.
Horner, J. R. 1979a. Upper Cretaceous dinosaurs from the Bearpaw Shale (marine) of south-central Montana with a checklist of Upper Cretaceous dinosaur remains from marine sediments in North America. Journal of Paleontology 53(3):566–577.
Horner, J. R. and Makela, R. 1979. Nest of juveniles provides evidence of family structure among dinosaurs. Nature 282:296–298.
Horner, J. R. 1982. Evidence of colonial nesting and 'site fidelity' among ornithischian dinosaurs. Nature 297:675–676.
Horner, J. R. 1983. Cranial osteology and morphology of the type specimen of Maiasaura peeblesorum (Ornithischia: Hadrosauridae), with discussion of its phylogenetic position. Journal of Vertebrate Paleontology 3(1):29–38.
Horner, J. R. 1984a. A "segmented" epidermal tail frill in a species of hadrosaurian dinosaur. Journal of Paleontology 58(1):270–271.
Horner, J. R. 1984c. Three ecologically distinct vertebrate faunal communities from the Late Cretaceous Two Medicine Formation of Montana, with discussion of evolutionary pressures induced by seaway fluctuations. Montana Geological Society Guidebook 1984:299–303.
Horner, J. R. 1985. Stratigraphic position of the Bear Gulch Limestone (Lower Carboniferous) of central Montana. C.r. IX Congr. Int. Strat. Geol. Carbonifére 5:427–436.
Horner, J. R. and Lund, R. 1985. Biotic distribution and diversity in the Bear Gulch Limestone of central Montana. C.r. IX Congr. Int. Strat. Geol. Carbonifére 5:437–442.
Weishampel, D. B. and Horner, J. R. 1986. The hadrosaurid dinosaurs from the Iren Dabasu fauna (Peoples Republic of China, Late Cretaceous). Journal of Vertebrate Paleontology 6(1):38–45.
Horner, J. R. 1987. "Ecological and behavioral implications derived from a dinosaur nesting site"; pp. 51–63 in S. J. Czerkas and E. C. Olson (eds.), Dinosaurs Past and Present, Vol. II. University of Washington Press, Seattle, Washington.
Weishampel, D. B. and Horner, J. R. 1987. "Dinosaurs, habitat bottlenecks, and the St. Mary River Formation"; pp. 224–229 in P. Currie and E. H. Koster (eds.), Fourth Symposium on Mesozoic Terrestrial Ecosystems. Occasional Paper of the Tyrrell Museum of Palaeontology, Drumheller, Alberta, Canada.
Horner, J. R. 1988b. A new hadrosaur (Reptilia: Ornithischia) from the Upper Cretaceous Judith River Formation of Montana. Journal of Vertebrate Paleontology 8(3):314–321.
Horner, J. R. and Weishampel, D. B. 1988. A comparative embryological study of two ornithischian dinosaurs. Nature 332:256–257.
Horner, J. R. 1989a. The Mesozoic terrestrial ecosystems of Montana. Montana Geological Society Guidebook 1989:153–162.
Horner, J. R. 1990. "Evidence of diphyletic origination of the hadrosaurian (Reptilia: Ornithischia) Dinosaurs" - Chap. 13; pp. 179–187 in K. Carpenter and P. Currie (eds.), Dinosaur Systematics: Perspectives and Approaches. Cambridge University Press, New York, New York.
Weishampel, D. B. and Horner, J. R. 1990. "Hadrosauridae" - Chap. 26; pp. 534–561 in D. B. Weishampel, P. Dodson and H. Osmolska (eds.), The Dinosauria. University of California Press, Berkeley, California.
Horner, J. R. 1992a. Cranial morphology of Prosaurolophus (Ornithischia: Hadrosauridae) with descriptions of two new hadrosaurid species and an evaluation of hadrosaurid phylogenetic relationships. Museum of the Rockies Occasional Paper #2:1–119, Bozeman, Montana.
Horner, J. R. Varricchio, D. J. and Goodwin, M. B. 1992. Marine transgressions and the evolution of Cretaceous dinosaurs. Nature 358:59–61.
Barreto, C., Albrecht, R. M., Bjorling, D. E., Horner, J. R. and Wilsman, N. J. 1993. Evidence of the growth plate and the growth of long bones in juvenile dinosaurs. Science 262:2020–2023.
Rogers, R. R., Swisher, C. C., Horner, J. R. 1993. Ar40/Ar39 age and correlation of the nonmarine Two Medicine Formation (Upper Cretaceous), northwestern Montana, USA. Canadian Journal of Earth Sciences 30:1066–1075.
Varricchio, D. J. and Horner, J. R. 1993. Hadrosaurid and Lambeosaurid bone beds from the Upper Cretaceous Two Medicine Formation of Montana: taphonomic and biologic implications. Canadian Journal of Earth Sciences 30:997–1006.
Horner, J. R. 1994. "Steak knives, beady eyes, and tiny little arms (A portrait of T-rex as a scavenger)"; pp. 157–164 in G. D. Rosenberg and D. L. Wolberg (eds.), Dino-Fest. The Paleontological Society, Special Bulletin 7.
Horner, J. R. 1994. "Comparative taphonomy of some dinosaur and extant bird colonial nesting grounds" – Chapter 8; pp. 117–121 in K. Carpenter, C. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies. Cambridge University Press, New York, New York.
Horner, J. R. and Currie, P. J. 1994. "Embryonic and neonatal morphology and ontogeny of a new species of Hypacrosaurus (Ornithischia: Lambeosauridae) from Montana and Alberta"; pp. 312–336 in K. Carpenter, C. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies. Cambridge University Press, New York, New York.
Maxwell, D. and Horner, J. R. 1994. Neonate dinosaurian remains and dinosaurian eggshell from the Cloverly Formation, Montana. Journal of Vertebrate Paleontology 14(1):143–146.
Weishampel, D. B. and Horner, J. R. 1994. "Life history syndromes, heterochrony, and the evolution of Dinosauria"; pp. 229–243 in K. Carpenter, C. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies. Cambridge University Press, New York, New York.
Padian, K., Ricqlès, A. de, and Horner, J. R. 1995. Bone histology determines identification of a new fossil taxon of pterosaur (Reptilia: Archosauria). Compte Rendue Academie Science Paris 320(2):77–84.
Horner, J. R. and Weishampel, D. B. 1996. Correction to: A comparative embryological study of two ornithischian dinosaurs (1988). Nature 383:103.
Ruben, J. A., Hillenius, W. J., Geist, N. R., Leitch, A., Jones, T. D., Currie, P. J., Horner, J. R. , and Espe III, G. 1996. The metabolic status of some Late Cretaceous dinosaurs. Science 273:1204–1207.
Horner, J. R. 1997. Rare preservation of an incompletely ossified fossil embryo. Journal of Vertebrate Paleontology 17(2):431–434.
Schweitzer, M. H., Johnson, C, Zocco, G., Horner, J. R. , and Starkey, J. 1997. Preservation of biomolecules in cancellous bone of Tyrannosaurus rex. Journal of Vertebrate Paleontology 17(2):349–359.
Schweitzer, M. H., Marshal, M., Carron, K., Bohle, D. S., Busse, S. C., Arnaold, E. V., Barnard, D., Horner, J. R., and Starkey, J. R. 1997. Heme compounds in dinosaur trabecular bone. Proceedings of the National Academy of Sciences 94:6291–6296.
Varricchio, D. J., Jackson, F., Borkowski,J., and Horner, J. R. 1997. Nest and egg clutches of the dinosaur Troodon formosus and the evolution of avian reproductive traits. Nature 385:247–249.
Horner, J. R. and Clouse, V. 1998. "An undisturbed clutch of hadrosaur eggs from the Judith River Formation of Montana"; pp. 22–25 in 1st International Meeting on Dinosaur Paleobiology, Museu Nacional de Historia Natural, Universidade de Lisboa, Spain.
Horner, J. R., Ricqlès, A. de, and Padian, K. 1999. Variation in dinosaur skeletochronology indicators: implications for age assessment and physiology. Paleobiology 25(4):295–304.
Horner, J. R. 1999. Egg clutches and embryos of two hadrosaurian dinosaurs. Journal of Vertebrate Paleontology 19(4):607–611.
Schweitzer, M. H. and Horner, J. R. 1999. Intravascular microstructures in trabecular bone of Tyrannosaurus rex. Ann. Paléontol. 85(3):179–192.
Horner, J. R. 2000. Dinosaur reproduction and parenting. Annual Reviews of Earth and Planetary Sciences 28:19–45.
Horner, J. R., Ricqlès, A. de, and Padian, K. 2000. Long bone histology of the hadrosaurid Maiasaura peeblesorum: growth dynamics and physiology based on an ontogenetic series of skeletal elements. Journal of Vertebrate Paleontology 20(1):109–123.
Ricqlès, A. de, Padian, K., Horner, J. R., and Francillon-Vieillot, H. 2000. Paleohistology of the bones of pterosaurs (Reptilia: Archosauria): anatomy, ontogeny, and biomechanical implications. Zoological Journal of the Linnean Society 129:349–385.
Horner, J. R., Padian, K., and de Ricqlès, A. 2001. Comparative osteohistology of some embryonic and perinatal archosaurs: developmental and behavioral implications for dinosaurs. Paleobiology 27(1):39–58.
Horner, J. R. 2001. "Dinosaur Ethology"; pp. 416–419 in Briggs and Crowther (eds.), Palaeobiology II. Blackwell Science, Oxford, England.
Horner, J. R. 2001. "Dinosaur Behaviour"; A3318 in Encyclopedia of Life Sciences. Macmillan References Ltd.
Horner, J. R., Schmitt, J., Jackson, F., and Hannah, R. 2001. "Bones and rocks of the Two Medicine/Judith River clastic wedge complex of Montana"; pp. 3–13 in C.L. Hill (ed.), SVP Field Guidebook. Museum of the Rockies Occasional Paper #3, Bozeman, Montana.
Padian, K., A. J. de Ricqlès, and J. R. Horner. 2001. Dinosaurian growth rates and bird origins. Nature 412:405–408.
Rayfield, E. J., Norman, D. B., Horner, C. C., Horner, J. R., Smith, P. M. Thompson, J. and Upchurch, P. 2001. Cranial design and function in a large theropod dinosaur. Nature 409:1033–1037.
Ricqlès, A. de, Padian, K., and Horner, J. R. 2001. "The bone histology of basal birds in phylogenetic and ontogenetic perspectives"; pp. 411–426 in Jacques Gauthier and Lawrence F. Gall (eds.), New Perspectives on the Origin and Early Evolution of Birds. Peabody Museum of Natural History, Yale University, New Haven, Connecticut.
Horner, J.R. 2002. "Evidence of dinosaur social behavior" - Chapter 9; pp. 71–78 in J. G. Scotchmoor, B. B. Breithaupt, D. A. Springer & T. R. Fiorillo (eds.), Dinosaurs, The Science behind the Stories. American Geological Institute, Alexandria, Virginia.
Padian, K. and Horner, J. R. 2002. Typology versus transformation in the origin of birds. Trends in Ecology and Evolution 17(3):120–124.
Varricchio, D. J., Horner, J. R. and Jackson, F. 2002. Embryos and eggs for the Cretaceous theropod Troodon formosus. Journal of Vertebrate Paleontology 22(3):564–576.
Chin, K., Eberth, D.A., Schweitzer, M.H., Rando, T.A., Sloboda, W.J., and Horner, J.R. 2003. Remarkable preservation of muscle tissue within a tyrannosaurid coprolite from Alberta, Canada. Palaios 18:286–294.
Ricqlès, A. de, Padian, K. and Horner, J. R. 2003. On the bone histology of some Triassic pseudosuchian archosaurs and related taxa. Annales de Palèontologie 89:67–101.
Ricqlès, A. de, Padian, K., Horner, J. R. , Lamm, E.-T., and Myhrvold, N. 2003. Osteohistology of Confuciousornis sanctus (Theropoda: Aves). Journal of Vertebrate Paleontology 23(2):373–386.
Goodwin, M. and Horner, J. R. 2004. Cranial histology of pachycephalosaurs (Ornithischia: Marginocephalia) reveals transitory structures inconsistent with head-butting behavior. Paleobiology 30(2):253–267.
Horner, J. R. and Padian, K. 2004. Age and growth dynamics of Tyrannosaurus rex. Proceedings Royal Society (B) 271(1551):1875–1880.
Horner, J. R., Weishampel, D. B., and Forster, C. 2004. "Hadrosauridae"; pp. 438–463 in Weishampel, Dodson and Osmolska (eds.), The Dinosauria. University of California Press, Berkeley, California.
Padian, K. and Horner, J.R. 2004. "Dinosaur physiology"; pp. 660–671 in Weishampel, Dodson and Osmolska (eds.), The Dinosauria. University of California Press, Berkeley, California.
Padian, K., Horner, J. R. and de Ricqlés, A. 2004. Little dinosaurs and pterosaurs; the evolution of archosaurian growth strategies. Journal of Vertebrate Paleontology 24:555–571.
Channing, A. C., Schweitzer, M. H., Horner, J. R., and McEneaney, T. 2005. A silicified bird from Quaternary hot spring deposits. Proceedings of the Royal Society (B) 272:905–911.
Main, R. P, Ricqlès, A. de, Horner, J. R., and Padian, K. 2005. The evolution and function of thyreophoran dinosaur scutes: implications for plate function in stegosaurs. Paleobiology 31(2):291–314.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2005. Gender-specific reproductive tissue in ratites and Tyrannosaurus rex. Science 308:1456–1460.
Schweitzer, M. H., Wittmeyer, J. L., Horner, J. R., and Toporski, J.B. 2005. Soft-tissue vessels and cellular preservation in Tyrannosaurus rex. Science 307:1952–1955.
Goodwin, M., Clemmens, B., Horner, J. R., and Padian, K. 2006. The smallest known Triceratops skull: new observations on ceratopsid cranial anatomy and ontogeny. Journal of Vertebrate Paleontology 26(1):103–112.
Horner, J. R. and Goodwin, M. 2006. Major cranial changes in Triceratops. Proceedings of the Royal Society (B) 273:2757–2761.
Prieto-Marquez, A., Weishampel, D. B., and Horner, J. R. 2006. The dinosaur Hadrosaurus foulkii, from the Campanian of the East Coast of North America, with a reevaluation of the genus. Acta Palaeontol. Pol. 51(1):77–98.
Ricqlès, A. de, Horner, J. R., and Padian, K. 2006. The interpretation of dinosaur growth patterns. Trends in Ecology and Evolution 21(11):596–597.
Chinnery, B. and Horner, J. R. 2007. A new neoceratopsian dinosaur linking North American and Asian taxa. Journal of Vertebrate Paleontology 27(3):625–641.
Lipkin, C., Sereno, P. C., and Horner, J. R. 2007. The furcula in Suchomimus tenerensis and Tyrannosaurus rex (Dinosauria: Theropoda: Tetanurae). Journal of Paleontology 81(6):1523–1527.
Schweitzer, M. H., Elsey, R. M., Dacke, C. G., Horner, J. R., and Lamm, E.-T. 2007. Do egg-laying crocodilian (Alligator mississippiensis) archosaurs form medullary bone? Bone 40:1152–1158.
Schweitzer, M. H., Suo, Z., Avci, R., Asara, J. M., Allen, M. A. Arce, F. T., and Horner, J. R. 2007. Analysis of soft tissue from Tyrannosaurus rex suggest the presence of protein. Science 316:277–280.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2007. Soft tissue and cellular preservation in vertebrate skeletal elements from the Cretaceous to the present. Proceedings of the Royal Society (B) 274:183–197.
Cooper, L. N., Lee, A. H., Taper, M. L., and Horner, J. R. 2008. Relative growth rates of predator and prey dinosaurs reflect effects of predation. Proc. Royal Soc. B 275:2609–2615.
Horner, J. R. and Goodwin, M. 2008. Ontogeny of cranial epi-ossifications in Triceratops. Journal of Vertebrate Paleontology 28(1):134–144.
Ricqlès, A. de, Padian, K., Knoll, F., and Horner, J. R., 2008. On the origin of high growth rates in archosaurs and their ancient relatives: Complementary histological studies on Triassic archosauriformes and the problem of a phylogenetic signal in bone histology. Annales de Paléontologie 94:57–76.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2008. "One pretty amazing T. rex." in Larson and Carpenter (eds.), Tyrannosaurus rex, The Tyrant King. Indiana University Press, Bloomington, Indiana.
Horner, J. R. and Goodwin, M. B. 2009. Extreme cranial ontogeny in the Upper Cretaceous dinosaur Pachycephalosaurus. PLoS ONE 4(10):e7626
Horner, J. R., Ricqlès, A. de, Padian, K., and Scheetz, R. 2009. Comparative long bone histology of the “hypsilophodontid” dinosaurs Orodromeus makelai, Dryosaurus altus, and Tenontosaurus tillettii (Ornithischia: Euornithopoda). Journal of Vertebrate Paleontology 29(3):1–14.
Schweitzer, M. H., Zheng, W., Organ, C. L., Avci, R., Suo, Z., Freimark, L. M., Lebleu, V. S., Duncan, M. B., Vander Heiden, M. G., Neveu, J. M., Lane, W. S., Cottrell, J. S., Horner, J. R., Cantly, L. C., Kalluri, R., and Asara, J. M. 2009. Biomolecular characterization and protein sequences of the Campanian hadrosaurs B. Canadensis. Science 324:626–631.
Wolff, E. D. S., Salisbury, S. W., Horner, J. R., and Varricchio, D. J. 2009. Common avian infection plagued the tyrant dinosaurs. PLoS ONE 4(9):e7288.
Archibald, J. D., et al. (29 authors). 2010. Cretaceous extinctions: Multiple causes. Science 328:973
Goodwin, M. B. and Horner, J. R. 2010. "Historical collecting bias and the fossil record of Triceratops in Montana"; pp. 551–563 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs. The Royal Tyrrell Museum Ceratopsian Symposium, Indiana University Press, Bloomington, Indiana.
Jackson, F. D., Horner, J. R., and Varricchio, D. J. 2010. A study of a Troodon egg containing embryonic remain using epifluorescence microscopy and other techniques. Cretaceous Research 31:255–262.
Longrich, N. R., Horner, J. R., Erickson, G. M., and Currie, P. J. 2010. Cannibalism in Tyrannosaurus rex. PLoS ONE 5(10):e13419.
McDonald, A. T. and Horner, J. R. 2010. "New material of “Styracosaurus” ovatus from the Two Medicine Formation of Montana." pp. 156–168 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs. The Royal Tyrrell Museum Ceratopsian Symposium, Indiana University Press, Bloomington, Indiana.
Scannella, J. B. and Horner, J. R. 2010. Torosaurus Marsh, 1891, is Triceratops Marsh, 1889 (Certatopisdae: Chasmosaurinae): synonomy through ontogeny. Journal of Vertebrate Paleontology 30(4):1157–1168.
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Therrien, F., and Horner, J. R. 2011. A Subadult skull of Hypacrosaurus stebingeri (Ornithischia: Lambeosaurinae): anatomy and comparison. Historical Biology 23(1):63–72.
Horner, J. R., Goodwin, M., and Myhrvold, N. 2011. Dinosaur census reveals abundant Tyrannosaurus and rare ontogenetic stages in the Upper Cretaceous Hell Creek Formation (Maastrichtian), Montana, USA. PLoS ONE 6(2):e16574.
Horner, J. R. and Lamm, E-T. 2011. Ontogeny of the parietal frill of Triceratops: a preliminary histological analysis. Comptes Rendus Palevol 10:439–452.
Fowler, D. W., Woodward, H. N., Freedman, E. A., Larson, P. L., and Horner, J. R. 2011. Reanalysis of “Raptorex kriegsteini”: a juvenile tyrannosaurid dinosaur from Mongolia. PLoS ONE 6(6):e21376.
Gates, T. A., Horner, J. R., Hanna, R. R., and Nelson, C. R. 2011. New unadorned hadrosaurine hadrosaurid (Dinosauria, Ornithopoda) from the Campanian of North America. Journal of Vertebrate Paleontology 31(4):798–811.
Padian, K. and Horner, J. R. 2011. The evolution of ‘Bizarre structures’ in dinosaurs: biomechanics, sexual selection, social selection or species recognition? Journal of Zoology 283(1):3–17. doi:10.1111/j.1469-7998.2010.00719.x
Padian, K. and Horner, J. R. 2011. The definition of sexual selection and its implications for dinosaurian biology. Journal of Zoology. 283(1):23–27. doi:10.1111/j.1469-7998.2010.00761.x
Prieto-Marquez, A., Minjin, B., and Horner, J. R. 2011. A diminutive deinonychosaur (Dinopsauria: Theropoda) from the Early Cretaceous of Oosh (Ovorkhangai, Mongolia). Alcheringa 36(1):117-136. doi:10.1080/03115518.2011.590401
Scannella, J. B. and Horner, J. R. 2011. ‘Nedoceratops’: An example of a transitional morphology. PLoS ONE 6(12):e28705. doi:10.1371/journal.pone.0028705
Schott, R. K., Evans, D. C., Goodwin, M. B. Horner, J. R., Brown, C. M., and Longrich, N. R. 2011. Cranial ontogeny in Stegoceras validum (Dinosauria: Pachycephalosauria): a quantitative model of pachycephalosaur dome growth and variation. PLoS ONE 6(6):e21092. doi:10.1371/journal.pone.0021092
Woodward, H. N., Horner, J. R., and Farlow, J. O. 2011. Osteohistological evidence for determinate growth in the American Alligator. Journal of Herpetology 45(3):339–342.
Bailleul, A. M., Hall, B. K., and Horner, J. R. 2012. First evidence of dinosaurian secondary cartilage in the post-hatching skull of Hypacrosaurus stebingeri (Dinosauria, Ornithischia). PLoS ONE 7(4):e36112. doi:10.1371/journal.pone.0036112
Horner, J. R. 2012. Dinosaur behaviour. in ELS. John Wiley & Sons, Ltd: Chickester. doi:10.1002/9780470015902.a0003318.pub3
Zelenitsky, D. K., Horner, J. R., and Therrien, F. 2012. Dinosaur Eggs: pp. 613–620 in M. K. Brett-Surman, T. R. Holtz and J.O. Farlow (eds.), The Complete Dinosaur. Indiana University Press, Bloomington, Indiana.
Padian, K. and Horner, J. R. 2013. The species recognition hypothesis explains exaggerated structures in non-avialan dinosaurs better than sexual selection does. Comptes Rendus Palevol 13(2): doi:10.1016/j.crpv.2013.10.004
Bailleul, A. M., Hall, B. K., Horner, J. R. 2013. Secondary cartilage revealed in a non-avian dinosaur embryo. PLoS ONE 8(2):e56937. doi:10.1371/journal.pone.0056937
Padian, K. and Horner, J. R. 2013. Misconceptions of sexual selection and species recognition: a response to Knell et al. and to Mendelson and Shaw. Trends in Ecology & Evolution 28(5): doi:10.1016/j.tree.2013.01.011
Goodwin, M. B. and Horner, J. R. 2014. Cranial morphology of a juvenile Triceratops skull from the Hell Creek Formation, McCone County, Montana, with comments on the fossil record of ontogenetically younger skulls. Special Paper of the Geological Society of America 503:333-347. doi:10.1130/2014.2503(13)
Horner, J. R. 2014. Paleontology: a cock's comb on a duck-billed dinosaur. Current Biology 24(2):R85-6. doi:10.1016/j.cub.2013.12.007
Woodward, H. N., Horner, J. R., Farlow, J. O. 2014. Quantification of intraskeletal histovariability in Alligator mississippiensis and implication for vertebrate osteohistology. PeerJ 2:e422. doi:10.7717/peerj.422
Scannella, J. B., Fowler, D. W., Goodwin, M. B., Horner, J. R. 2014. Evolutionary trends in Triceratops from the Hell Creek Formation, Montana. Proceedings of the National Academy of Sciences 111(28):10245-10250. doi:10.1073/pnas.1313334111
Rashid, D. J., Chapman, S. C., Larsson, H. C. E., Organ, C. L., Bebin, A. G., Merzdorf, C. S., Bradley, R., Horner, J. R. 2014. From dinosaurs to birds: a tail of evolution. EvoDevo 5(25):1-20. doi:10.1186/2041-9139-5-25
Padian, K. and Horner, J. R. 2014. Darwin's sexual selection: understanding his ideas in context. Comptes Rendus Palevol 13(8). doi:10.1016/j.crpv.2014.09.001
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Horner, J. R., Therrien, F. 2014. Cranial morphology and variation in Hypacrosaurus stebingeri (Ornithischia: Hadrosauridae): chapter 14: pages 245-276 in Eberth, D. A. and Evans, D. C (eds.). Hadrosaurs. Indiana University Press, Bloomington, Indiana.
Bailleul, A. M., Nyssen-Behets, C., Lengele, B., Hall, B. K., Horner, J. R. 2015. Chondroid bone in dinosaur embryos and nestlings (Ornithischia: Hadrosauridae): insights into the growth of the skull and the evolution of skeletal tissues. Comptes Rendus Palevol (in press). doi:10.1016/j.crpv.2014.12.004
Horner, J. R., Woodward, H. N., Bailleul, A. M. 2015. Mineralized tissues in dinosaurs interpreted as having formed through metaplasia: a preliminary evaluation. Comptes Rendus Palevol (in press). doi:10.1016/j.crpv.2015.01.006
Padian, K., Werning, S., Horner, J. R. 2015. A hypothesis of differential secondary bone formation in dinosaurs. Comptes Rendus Palevol
(in press). doi:10.1016/j.crpv.2015.03.002
Cubo, J., Woodward, H. N., Wolff, E., Horner, J. R. 2015. First reported cases of biomechanically adaptive bone modeling in non-avian dinosaurs. PLoS ONE 10(7). doi:10.1371/journal.pone.0131131
Published Abstracts (102):
Baird, D. and Horner, J. R. 1977. A fresh look at the dinosaurs of New Jersey and Delaware. New Jersey Acad. Sci. Bulletin 22(2):50.
Lund, R. and Horner, J. R. 1979. Biotic distribution and diversity in the Bear Gulch Basin. Ninth International Congress of Carboniferous Stratigraphy and Geology Abstracts:120.
Melton, W. G., Lund, R. and Horner, J. R. 1979. Faunal distribution within the Bear Gulch lentil. Ninth International Congress of Carboniferous Stratigraphy and Geology Abstracts:136.
Horner, J. R. 1987. Evolution of terrestrial Late Cretaceous faunal communities. Journal of Vertebrate Paleontology 7(3, Supplement):18A.
Currie, P. J. and Horner, J. R. 1988. Lambeosaurine hadrosaur embryos (Reptilia: Ornithischia). Journal of Vertebrate Paleontology 8 (3, Supplement):13A.
Horner, J. R. 1988a. Morphological evidence of mammalian-grade herd structure in hadrosaurian and ceratopsian dinosaurs. International Symposium on Vertebrate Behavior as Derived from the Fossil Record 1988:13. Montana State University, Bozeman, Montana.
Horner, J. R. 1989b. The ontogenetic predictability of hadrosaur crania. Journal of Vertebrate Paleontology 9(3, Supplement):25A.
Horner, J. R. 1988c. Cranial allometry of Maiasaura peeblesorum (Ornithischia; Hadrosauridae) and its behavioral significance. Journal of Vertebrate Paleontology 8(3, Supplement):18A.
Cooley, J., Schmitt, J., and Horner, J. R. 1992. Fluvial systems of the Upper Jurassic Morrison Formation, Gallatin and Beartooth Ranges, SW Montana. SEPM Annual Meeting Bulletin 1992.
Horner, J. R. 1992b. Dinosaur behavior and growth. Paleontological Society Special Publication 6:135.
Barreto, C. B., Albrecht, R. M., Horner, J. R., and Wilsman, N. H. 1993. Evidence of dinosaur growth plate morphology in the fossil record. The Society for Orthopedic Research, Annual Meeting 1993.
Barreto, C. B., Albrecht, R. M., Horner, J. R., and Wilsman, N. H. 1993. Dinosaur long bone growth plates. Journal of Vertebrate Paleontology 13(3, Supplement):25A.
Clouse, V. and Horner, J. R. 1993. Eggs and embryos from the Judith River Formation of Montana. Journal of Vertebrate Paleontology 13(3, Supplement):42A.
Horner, J. R. 1993. Ontogenetic development of the lambeosaurid narial crest. Journal of Vertebrate Paleontology 13(3, Supplement):42A.
Padian, K., Horner, J. R., and de Ricqlès, A. 1993. A new azhdarchid pterosaur from the Two Medicine Formation (Late Cretaceous: Campanian) of Montana. Journal of Vertebrate Paleontology 13(3, Supplement):52A
Ricqlès, A. de, Padian, K., and Horner, J. R. 1993. Paleohistology of pterosaur bones. Journal of Vertebrate Paleontology 13(3, Supplement):54A.
Schweitzer, M. H., Cano, R. J., Starkey, J. R., and Horner, J. R. 1994. Multiple lines of evidence for the preservation of collagen and other biomolecules in unmineralized bone from Tyrannosaurus rex. Journal of Vertebrate Paleontology 14(3, Supplement):45A.
Carter, D. R., Mikic, B., and Horner, J. R. 1995.Growth plate differences between mammals and dinosaurs are related to both genotypic and mechanical factors. Journal of Vertebrate Paleontology 15(3, Supplement):22A
Horner, J. R. 1995. Morphology and function of the enclosed narial chambers of lambeosaurid dinosaurs. Journal of Vertebrate Paleontology 15(3, Supplement):36A.
Horner, J. R. 1996a. Transgressions, regressions, and the evolution of dinosaurs. The Paleontological Society, Special Publication 8:178.
Horner, J. R. 1996b. New evidence for post-eclosion parental attention in Maiasaura peeblesorum. Journal of Vertebrate Paleontology 16(3, Supplement):42A.
Laws, R., Hasiotis, S. T., Fiorillo, A. R., Chure, D., Breithaupt, B. H., and Horner, J. R. 1996. The demise of a Jurassic Morrison dinosaur after death - three cheers for the dermestid beetle. Geological Society of America, Abstracts with Programs 28:530A.
Goodwin, M. B., Clemens, W. A., and Horner, J. R. 1997. Morphological variation and ontogeny in the skull of Triceratops. Journal of Vertebrate Paleontology 17(3, Supplement):49A.
Horner, J. R., Ricqlès, A. de, and Padian, K. 1997. Histological analysis of a dinosaur skeleton: evidence of skeletal growth variation. Journal of Morphology 232(3):267.
Horner, J. R. and Varricchio, D. J. 1997. Embryonic remains of Troodon formosus from the Two Medicine Formation of Montana. Journal of Vertebrate Paleontology 17(3, Supplement):53A.
Maxwell, W., Hallas, B. H., and Horner, J. R. 1997. Further neonate dinosaurian remains and dinosaurian eggshell from the Cloverly Formation of Montana. Journal of Vertebrate Paleontology 17(3):63A.
Padian, K., Ricqlès, A. de, and Horner, J. R. 1997. Pterosaur bone histology: growth rates and mechanics. Journal of Morphology 232(3):306.
Ricqlès, A. de, Horner, J. R., and Padian, K. 1997. Histological evidence of dinosaur growth patterns. Abstracts, Third World Herpetological Congress, Prague.
Ricqlès, A. de, Padian, K., and Horner, J. R. 1997. Comparative biology and the bone histology of extinct tetrapods: what does it tell us? Journal of Morphology 232(3):246.
Varricchio, D. J., Jackson, F., Borkowski, J. J., and Horner, J. R. 1997. Coelurosaurian eggs: between a rock and a soft place. Journal of Vertebrate Paleontology 17(3, Supplement):82A.
Ancell, C. A., Harmon, R., and Horner, J. R. 1998. Gar in a duckbill: preservation of a scavenger and its prey. Journal of Vertebrate Paleontology 18(3, Supplement):24A
Cooper, L. N., Lamm, E., Horner, C. C. and Horner, J. R. 1998. Annual changes in mass of Hypacrosaurus stebingeri as hypothesized from lines of arrested growth. Journal of Vertebrate Paleontology 18(3, Supplement):39A
Horner, J. R. and Goodwin, M. B. 1998. Did pachycephalosaurs really head-butt? An osteohistological cranial analysis. Journal of Vertebrate Paleontology 18(3, Supplement):52A
Ricqlès, A. de, Horner, J. R., and Padian, K. 1998. Growth dynamics of the hadrosaurid dinosaur Maiasaura peeblesorum. Journal of Vertebrate Paleontology 18(3, Supplement):72A.
Schmitt, J. G., Horner, J. R., Laws, R. R. and Jackson, F. 1998. Debris-flow deposition of a hadrosaur-bearing bone-bed, Upper Cretaceous Two Medicine Formation, northwest Montana. Journal of Vertebrate Paleontology 18(3, Supplement):76A.
Cooper, L. N. and Horner, J. R. 1999. Growth rate of Hypacrosaurus stebingeri as hypothesized from lines of arrested growth and whole femur circumference. Journal of Vertebrate Paleontology 19(3, Supplement):39A.
Hanna, R. R., LaRock, J. W., and Horner, J. R. 1999. Pathological brachylophosaur bones from the Upper Cretaceous Judith River Formation, northeastern Montana. Journal of Vertebrate Paleontology 19(3, Supplement):49A.
Horner, J. R. 1999. Dinosaur biozones in the Campanian clastic wedge complex of Montana, and their biological implications. AAPG, Rocky Mountain Section, Abstracts with Program 1999:A6.
Horner, J. R., Padian, K., and de Ricqlès, A. 1999. Osteohistology of some embryonic and perinatal archosaurs: phylogenetic and behavioral implications for dinosaurs. Journal of Vertebrate Paleontology 19(3, Supplement):53A.
Padian, K., Horner, J. R., and de Ricqlès, A. 1999. Dinosaurian growth rates and the evolution of life history strategies. Journal of Vertebrate Paleontology 19(3, Supplement):67A.
Ricqlès, A. de, Padian, K., and Horner, J. R. 1999. The bone histology of basal birds in phylogenetic and ontogenetic perspectives. Journal of Vertebrate Paleontology 19(3, Supplement):70A.
Main, R. P., Padian, K, and Horner, J. R. 2000. Comparative histology, growth, and evolution of archosaurian osteoderms: why did Stegosaurus have large dorsal plates? Journal of Vertebrate Paleontology 20(3):56A.
Barreto, C., Marshall, C., and Horner, J. R. 2001. Differential and determinate growth in dinosaurs and other reptiles. Journal of Vertebrate Paleontology 21(3):31A.
Goodwin, M. and Horner, J. R. 2001. How Triceratops got its horns: New information from a growth series on cranial morphology and ontogeny. Journal of Vertebrate Paleontology 21(3):56A.
Higby Schweitzer, M., Schmitt, J.G., 2001, and Horner, J.R. 2001. Paleoenvironmental and taphonomic parameters of an exceptionally preserved Tyrannosaurus rex. Geological Society of America Abstracts 33(6):388.
Horner, C. C., Horner, J. R., and Weishampel, D. B. 2001.Comparative internal cranial morphology of some hadrosaurian dinosaurs using computerized tomographic X-ray analysis and rapid prototyping. Journal of Vertebrate Paleontology 21(3):64A
Horner, J. R. and Padian, K. 2001. Evolution of archosaurian growth strategies. Journal of Vertebrate Paleontology 21(3):63A
Jerison, H. J., Horner, J. R., and Horner, C. C. 2001. Dinosaur forebrains. Journal of Vertebrate Paleontology 21(3):65A.
Padian, K., Horner, J. R., de Ricqlès. 2001. Dinosaurian life history strategies: evidence of differential growth rates from bone histology. American Zoologist 40(5).
Ricqlès, A de, Padian, K., and Horner, J. R. 2001. Bone histology in extant and extinct archosauromorphs: A structurofunctional exploration in ontogeny and phylogeny. Journal of Morphology 248(3):222.
Horner, J. R. and Marshall, C. 2002. Keratin covered dinosaur skulls. Journal of Vertebrate Paleontology 22(3):67A.
Padian, K, Horner, J. R., and Ricqlès, A. 2002. How dinosaurs grew: growth rates and strategies. Geological Society of America.
Ricqlès, A. de, Padian, K., and Horner, J. R. 2002. On the bone histology of some Triassic pseudosuchian archosaurs and related taxa: a preliminary report. 8th International Symposium on Mesozoic Terrestrial Ecosystems, Cape Town, South Africa.
Chinnery, B. and Horner, J. R. 2003. New basal neoceratopsioan from the lower Two Medicine Formation of Montana provides a link between Asian and North America taxa. Journal of Vertebrate Paleontology 23(3):40A.
Prieto-Marquez, A., Weishampel, D. B., and Horner, J. R. 2003. Taxonomy and systematics of the holotype of Hadrosaurus foulkii (Dinosauria, Ornithopoda) from the Late Cretaceous of eastern North America. Journal of Vertebrate Paleontology 23(3):88A.
Schweitzer, M., Channing, A., Horner, J. R., and Chiappe, L. 2003. Microbial biomineralization and exceptional preservation of vertebrate fossils. Journal of Vertebrate Paleontology 23(3):95A.
Wolff, E. and Horner, J. R. 2003. The first occurrence of Orodromeus makelai postcrania from the Late Campanian Judith River Formation of eastern Montana. Journal of Vertebrate Paleontology 23(3):112A.
Horner, J. R. and Padian, K. 2004. Age and growth dynamics of Tyrannosaurus rex. Journal of Vertebrate Paleontology 24 (3):72A.
Padian, K., Horner, J. R., and Dhaliwal, J. 2004. Species recognition as the principal cause of bizarre structures in dinosaurs. Journal of Vertebrate Paleontology 23(3):100A.
Schweitzer, M., Wittmeyer, J., and Horner, J. R. 2004. A novel dinosaurian tissue exhibiting unusual preservation. Journal of Vertebrate Paleontology 24(3):111A.
Currie, P. J., Henderson, M., Horner, J. R., and Williams, S. A. 2005. On tyrannosaur teeth, tooth positions and the taxonomic status of Nanotyrannus lancensis. The Origin, Systematics, and Paleobiology of Tyrannosauridae Symposium Abstracts:19. Burpee Museum, Rockford, Illinois
Horner, J. R. and Goodwin, M. 2005. A new Triceratops cranial growth series. Journal of Vertebrate Paleontology 25(3):71A.
Padian, K., Horner, J. R., and Lee, A. 2005. Sexual dimorphism in dinosaurs? A review of the evidence. Journal of Vertebrate Paleontology 25(3):98A.
Schweitzer, M. H., Wittmeyer, J. L., and Horner, J. R. 2005. Reproductive tissues in exceptionally well preserved Tyrannosaurus rex (MOR 1125). The Origin, Systematics, and Paleobiology of Tyrannosauridae Symposium Abstracts:30, Burpee Museum, Rockford, Illinois.
Horner, J. R. 2006. Osteohistology in dinosaur research. AAAS symposium, St. Louis, MO.
Padian, K., de Ricqlès, A, and Horner, J. R. 2006. Comparative long bone histology and growth of some hypsilophodontid dinosaurs. Journal of Vertebrate Paleontology 26(3):108A.
Reizer, J, and Horner, J. R. 2006. An ontogenetic series of the ceratopsid dinosaur Einiosaurus procurvicornis as determined by long bone histology. Journal of Vertebrate Paleontology 26(3):114A.
Ricqlès, A. de, Padian, K., and Horner, J. R. 2006. Complementary studies on archosauriform and basal archosaur bone histology: is research on a plesiomorphic histological condition warranted? Chinese Symposium.
Zelenitsky, D., Therrien, F., Currie, P., Henderson, D., and Horner, J. R. 2006. Locomotory behavior in the lambeosaurine Hypacrosaurus stebingeri. Journal of Vertebrate Paleontology 26(3):143A.
DeBoef, M., Larsson, H., and Horner, J. R. 2007. Measurements of vasculature in Tyrannosaurus rex show a relationship between growth rate and vascular orientation. Journal of Vertebrate Paleontology 27(3):76A.
Horner, J. R., Goodwin, M. B., and Woodward, H. 2007. Synonymy consequences of dinosaur cranial ontogeny. Journal of Vertebrate Paleontology 27(3):92A.
Brink, K., Zelenitsky, D., Evans, D., and Horner, J.R. 2008. New subadult Hypacrosaurus stebingeri skulls and a comparative analysis of allometric crest growth in lambeosaurines. Journal of Vertebrate Paleontology 28(3):56A.
Evans, D., Witmer, L., Ridgely, R., and Horner, J. R. 2008. Endocranial anatomy of lambeosaurine dinosaurs: implications for cranial crest function and evolution. Journal of Vertebrate Paleontology 28(3):75A.
Lee, A., Cooper, L., Taper, M., and Horner, J. R. 2008. Rapid growth of the hadrosaur Hypacrosaurus reflects direct and indirect effects of predation. Journal of Vertebrate Paleontology 28(3):104A.
Horner, J. R. 2008. Dinosaur Cranial Ontogeny. Paleofest, Burpee Museum, Rockford, Illinois.
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Horner, J. R., and Therrien, F. 2009. Ontogenetic development of the skull of Hypacrosaurus stebingeri. Fourth International Symposium on Dinosaur Eggs and Babies. Museum of the Rockies Occasional Paper #4:12, Bozeman, Montana.
Goodwin, M. B. and Horner, J. R. 2009. The fossil record of Triceratops from the Hell Creek Formation, Montana. pp. 147 in 9th North American Paleontological Convention, Abstracts. Cincinnati Museum Center Scientific Contributions #3, Cincinnati, Ohio.
Horner, J. R., Goodwin, M. B. and Myhrvold, N. 2009. Hell Creek Formation dinosaur census reveals abundant Tyrannosaurus. pp. 146 in 9th North American Paleontological Convention, Abstracts. Cincinnati Museum Center Scientific Contributions #3, Cincinnati, Ohio.
Horner, J. R. and Lamm, E-T. 2009. Metaplasia provided dinosaur skulls extreme morphological plasticity during ontogeny. Journal of Vertebrate Paleontology 29(3):117A.
Jackson, F. D., Horner, J. R., and Varricchio, D. 2009. A study of a Troodon egg containing embryonic remains using epiflourescence microscopy and other techniques. Fourth International Symposium on Dinosaur Eggs and Babies. Museum of the Rockies Occasional Paper #4:20, Bozeman, Montana.
Jackson, F. D., Horner, J. R., and Varricchio, D. 2009. A study of a Troodon egg containing embryonic remains using epiflourescence microscopy and other techniques. Journal of Vertebrate Paleontology 29(3):121A.
Padian, K., Horner, J. R., Fowler, D., and Scannella, J. 2010. How a synergy of species recognition and social signaling explains cranial anatomy and ontogeny in several groups of dinosaurs. Journal of Vertebrate Paleontology 30(4, Supplement):143A.
Woodward, H. and Horner, J. R. 2010. Osteohistological analysis of Alligator mississippiensis indicates absence of fibrolamellar bone in crocodilians and confirms determinate growth with the first report of external fundamental system: implications for tetrapod histology. Journal of Vertebrate Paleontology 30(4, Supplement):190A.
Baillieu, A. and Horner, J. R. 2011. Early ossification and calcified tissues in the skull of Hypacrosaurus stebingeri (Ornithischia, Lambeosaurinae): a preliminary study. pp. 11 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology, Drumheller, Alberta, Canada.
Bailleul, A. M. and Horner, J. R. 2011. Perinatal histomorphogenesis of craniofacial elements of Hypacrosaurus stebingeri (Ornithischia, Lambeosauridae): a preliminary study. Vol.1:pp. 57 in International Symposium on Paleohistology, Abstracts (M. Kohler, X. Jordana and N. M. Moratalla Eds), Institut Català de Paleontologia, Barcelona, Spain.
Bailleul, A. and Horner, J. R. 2011. Early ossification and calcified tissues in the skull of Hypacrosaurus stebingeri (Ornithischia, Lambeosaurinae): a preliminary study. Journal of Vertebrate Paleontology 31(5, abstracts vol):65.
Baker, K., Scannella, J., Hall, L., and Horner, J. R. 2011. Biogeographic implications of a partial ceratopsid skeleton from the lower Two Medicine Formation (Campanian), Montana. Journal of Vertebrate Paleontology 31(5, abstracts vol):65.
Brink, K. S., Zelenitsky, D. K., Evans, D. C., Horner, J. R., and Therrien, F. 2011. Hypacrosaurus stebingeri: an exceptional record of dinosaurian cranial ontogeny. pp. 14-15 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology.
Evans, D. C., Currie, P. J., Witmer, L. M., and Horner, J. R., and Ebreth, D. 2011. A new low crested lambeosaurine hadrosaurid from the Dinossaur Park Formation of Sandy Point, eastern Alberta. pp. 48–49 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology, Drumheller, Alberta, Canada.
Fowler, D., Scannella, J., and Horner, J. R. 2011 Reassessing ceratopsid diversity using unified frames of reference. Journal of Vertebrate Paleontology 31(5, abstracts vol):111.
Goodwin, M., Stanton, K., Horner, J. R., and Carlson, S. 2011. Oxygen isotopic variability and preservation in Tyrannosaurus rex, modern ratites and crocodilians: revisiting the thermophysiology of T.rex using δ180. Journal of Vertebrate Paleontology 31(5, abstracts vol):117.
Horner, J. R. and Woodward, H. N. 2011. Riddle of the humongous hadrosaurs: what these giants reveal about dinosaur ontogeny, evolution, and ecology. Vol.1:pp. 66 in Hadrosaur Symposium Abstracts, Royal Tyrrell Museum of Paleontology, Drumheller, Alberta, Canada.
Horner, J. R., Woodward, H. N., and Bailleul, A. 2011. Evaluating maturity in dinosaurs. Vol.1:pp. 53 in International Symposium on Paleohistology, Abstracts (M. Kohler, X. Jordana and N. M. Moratalla Eds), Institut Català de Paleontologia, Barcelona, Spain.
Lawver, D., Jackson, F., and Horner, J. R. 2011. Turtle eggs and embryos in a clutch from the Upper Cretaceous (Campanian) Judith River Formation of Montana. Journal of Vertebrate Paleontology 31(5, abstracts vol):141.
Scannella, J., Fowler, D., Trevethan, I., Roberts, D., and Horner, J. R. 2011. Individual variation in Triceratops from the Hell Creek Formation, Montana: implications for dinosaur taxonomy. Journal of Vertebrate Paleontology 31(5, abstracts vol):187.
Woodward, H. N., and Horner, J. R. 2011. A robust single element microanalysis of a hadrosaurid dinosaur (Ornithopoda: Maiasaura peeblesorum) population reveals ontogenetic trends in growth and development. Vol.1:pp. 56 in International Symposium on Paleohistology, Abstracts (M.Kohler, X. Jordana and N. M. Moratalla Eds), Institut Català de Paleontologia, Barcelona, Spain.
Fowler, D. W., Scannella, J. B., Goodwin, M. G., and Horner, J. R. 2012. How to eat a Triceratops: large sample of toothmarks provides new insight into the feeding behavior of Tyrannosaurus. Journal of Vertebrate Paleontology 32(5, abstracts vol):96.
Horner, V. and Horner, J. R. 2012. Sturgeon dorsal osteoderm ontogeny: A transformational model for marginocephalian squamosal ornaments. Journal of Vertebrate Paleontology 32(5, abstracts vol):113.
Scannella, J. B., Fowler, D.W., Goodwin, M. G., and Horner, J. R. 2012. Transitional Triceratops: Details of an ontogenetic sequence from the upper Middle Unit of the Hell Creek Formation, Montana. Journal of Vertebrate Paleontology 32(5, abstracts vol):166.
Woodruff, C., and Horner, J. R. 2012. A re-evaluation of the vertebral Soft Tissue reconstruction within Dinosauria based on alternate extant analogues. Journal of Vertebrate Paleontology 32(5, abstracts vol):195.
Woodward, H.N., Horner J. R., and Farlow, J.O. 2012. Paleobiological implications of the growth history and histovariability in a population of the hadrosaurid dinosaur Maiasaura peeblesorum. Journal of Vertebrate Paleontology 32(5, abstracts vol):196.
Boatman, E. M., Goodwin, M. B., Holman, H., Fakra, S., Schweitzer, M. H., Gronsky, R., Horner, J. R. 2014. Synchrotron chemical and structural analysis of Tyrannosaurus rex blood vessels: the contribution of collagen hypercrosslinking to tissue longevity. Microscopy and Microanalysis 20(Suppl 3):1430-1431.
Edited Book (1):
Carpenter, K., Hirsch, K. F., and Horner, J. R. (eds.). 1994. Dinosaur Eggs and Babies, Cambridge University Press, New York, 372 pp.
Popular Books (8):
Horner, J. R. and J. Gorman. 1985. Maia, a Dinosaur Grows Up. Museum of the Rockies, Bozeman, Montana & Running Press, Philadelphia Pensylvannia, 34 pp.
Horner, J. R. and J. Gorman. 1988. Digging Dinosaurs. Workman Publishing, New York, 210 pp.
Horner, J. R. and D. Lessom. 1992. Digging up Tyrannosaurus rex. Crown Publishers, New York, 36 pp.
Horner, J. R. and D. Lessom. 1993. The Complete T-rex. Simon and Schuster, New York, 235 pp.
Horner, J. R. and E. Dobb. 1997. Dinosaur Lives. Harper Collins, New York, 244 pp.
Horner, J. R. 2001. Dinosaurs Under the Big Sky. Mountain Press, Missoula, Montana, 194 pp.
Horner, J. R. 2007. Digging Up Dinosaurs. Farcountry Press, Helena, Montana, 49pp.
Horner, J. R. and J. Gorman. 2009. How to Build a Dinosaur; Extinction Doesn’t Have to be Forever. Dutton, New York, 246 pp.
Popular Publications (46-Total # Articles)
Horner, J. R. 1982. "Coming home to roost." Montana Outdoors 13(4):2–5.
Horner, J. R. 1984. "The nesting behavior of dinosaurs." Scientific American 250(4):130–137.
Horner, J. R. and D. B. Weishampel. 1989. "Dinosaur eggs: the inside story." Natural History 89(12):60–67.
Horner, J. R. 1996-2001 (bi-monthly column) Wild Outdoor World, Rocky Mountain Elk Foundation, Children’s magazine, Montana.
Horner, J. R. 1997. "Behavior"; pp. 45–50 in P. Currie and K. Padian (eds.), The Encyclopedia of Dinosaurs. Academic Press, San Diego, California.
Horner, J. R. 1997. "Willow Creek Anticline"; pp. 786 in P. Currie and K. Padian (eds.), The Encyclopedia of Dinosaurs. Academic Press, San Diego, California.
Horner, J. R. 1998. "Dinosaur behavior." National Forum 78(3):12–15.
Horner, J. R. 2000. "Dinosaur." Encyclopedia Americana 9:123–131.
Horner, J. R. 2000. "Hadrosaur." Encyclopedia Americana ###
Horner, J. R. 2004. Jack Horner: "An Intellectual Autobiography." in Montana Professor , 14(2):15–19. University of Montana, Missoula, Montana.
Horner, J. R., Padian, K, and de Ricqlès, A. 2005. "How dinosaurs grew so large and so small." Scientific American 293(4):32–39.
Horner, J. R. 2005. "Dinosaur Behaviour", Encyclopedia of Life Sciences, John Wiley & Sons, Ltd.
Horner, J. R. 2007. "Discovery of Maiasaura." Cosmos Magazine.
Horner, J. R. 2008. "The extraordinary characteristics of dyslexia." Perspectives on Language and Literacy 34:35. International Dyslexia Association.
Horner, J. R. 2009. "Dinosaurs at the table" - Chapter 6; pp. 111–121 in D. Seplowski and M. Ruse (eds.) The Paleobiological Revolution; Essays on the Growth of Modern Paleontology. University of Chicago Press, Chicago, Illinois.
Horner, J. R. and J. Gorman. 2009. "Dinosaur Resurrection." Discover Magazine, April 2009:50–53.
Horner, J. R. 2012. "Why do animals that fly have feathers, not including bats"?; pp. 99–100 in Gemma Elwin Harris (ed.), Big Questions from Little People. Farber & Farber Lmt., London.
Horner, J. R. and Horner, V. S. 2012. "Time traveling in Montana’s Mesozoic park." in Distinctly Montana, Fall, 2012.
Horner, J. R. and Druckenmiller, P. 2012. "Polar Dinosaurs"; pp. 127–130 in D. S. Miller (ed.), On Arctic Ground. Braid River Press, Seattle, Washington.